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Red Deer stag (Cervus elaphus) in snow-covered pine forest, Cairngorms National Park, Scotland, UK. December
Red deer (Cervus elaphus) stag in the snow, Scotland, March
Edible frog (Rana esculenta) male calling at pond surface, Klein Schietveld, Brasschaat, Belgium May
Roe deer (Capreolus capreolus) stag peering through vegetation, Peerdsbos, Brasschaat, Belgium, July
Common Pheasant (Phasianus colchicus) male, Texel, the Netherlands, April
BlackbuckA(Antilope cervicapra), herd with males and females, Velavadar National ParkBlackbuck(Antilope cervicapra), herd with males and females, Velavadar National Park, Gujarat, India
Peacock (Pavo cristatus) on rock with Palm Squirrel (Funambulus sp) India
Adult bull African Elephant (Loxononta africana) reaching up to feed on foliage on the banks of the Luangwa River, South Luangwa National Park, Zambia, October
Victorias riflebird (Ptiloris victoriae) male displaying in rainforest, Atherton Tablelands, Queensland, Australia, October
Male Grizzly bear (Ursus arctos horribilis), Khutzeymateen Grizzly Bear Sanctuary, British Columbia, Canada, June
Male White breasted nuthatch (Sitta carolinensis) perched on stump, New York, USA, October
RF- Capercaillie (Tetrao urogallus) male displaying in heavy snowfall, Cairngorms National Park, Scotland. February
RF- Male Great bustard (Otis tarda) displaying, La Serena, Extremadura, Spain. April
RF- Red deer (Cervus elaphus) stag in pine forest in snow blizzard. Alvie Estate, Cairngorms NP, Highlands, Scotland, UK, March
Herd of Red deer (Cervus elaphus) stags in clearing in snow covered pine forest, Cairngorms National Park, Scotland, UK, December
Ptarmigan (Lagopus mutus) male in snowy mountains, Cairngorms National Park, Scotland, UK, May 2015
Black grouse (Tetrao tetrix) male displaying at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Osprey (Pandion haliaetus) male flying, Cairngorms National Park, Scotland, UK, May
Osprey (Pandion haliaetus) male flying with talons outstretched, Glenfeshie, Cairngorms National Park, Scotland, UK, May
Two Black grouse (tetrao tetrix) males fighting at lek, Deeside, Cairngorms National Park, Scotland, UK, April
Two Black grouse (Tetrao tetrix) males displaying at a lek site at dawn, Cairngorms National Park, Scotland, UK, April
Male impalas (Aepyceros melampus) standing in heavy rain, Masai-Mara Game Reserve, Kenya. February
Male black rhinoceros (Diceros bicornis) silhouetted against clouds, Masai-Mara Game Reserve, Kenya. March
Male Lion (Panthera leo) shaking off water in the rain, Masai-Mara game reserve, Kenya
Portrait of male Waterbuck (Kobus ellipsiprymnus) in the rain, Masai-Mara game reserve, Kenya
Pair of Rough snout ghostpipefish (Solenostomus paegnius), with female with egg pouch in front of male. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Portrait of male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, to feed on plankton. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Male Long tailed duck (Clangula hyemalis) on water, Batsfjord, Norway, March
Male King eider duck (Somateria spectabilis) on water, Batsfjord, Norway, March
Mallard (Anas platyrhynchos) walking on ice, Acadia National Park, Maine, USA, April
Two male Bearded reedling (Panurus biarmicus) eating seeds from the spike of a common bulrush (Typha latifolia) in a reedbed, Belgium, January. Digital composite
Brown hare (Lepus europaeus) two males chasing female in snow covered field during a blizzard. Derbyshire, England, UK, February. Winner of British Seasons category of BWPA competition 2014
Moorland Clouded Yellow butterfly (Colias palaeno) male and female, Joutsa (formerly Leivonmaki), Finland, July
Whooper Swan (Cygnus cygnus) male and female facing in opposite directions, central Finland, April
Black-and-white tuxedo male kitten, Tuxie, 8 weeks, with black-and-white Dutch rabbit, against white background
Fallow deer (Dama dama) male, Studen Kladenets reserve, Eastern Rhodope Mountains, Bulgaria, May 2013
Male green Darwins Frog (Rhinoderma darwinii) with young in vocal pouch, Chile, December 2012, Vulnerable species
A portrait of a large male Dusky grouper (Epinephelus marginatus). Lavezzi Islands, between Sardinia and Corsica in the Mediterranean Sea
Leopard (Panthera pardus) young male, portraits, Masai-Mara Game Reserve, Kenya
Two male Alpine ibex (Capra ibex ibex) fighting in front of the Mer de Glace glacier, with a female watching, Aiguilles Rouges (Red Peaks) Regional Natural Park, Haute-Savoie, France, June
Two male Black grouse (Lyrurus tetrix) displaying at lek, Utajarvi, Finland, April
Young roe deer buck (Capreolus capreolus) standing in snow, with tongue poking out. Southern Norway, February
Male Panther Chameleons (Furcifer pardalis) fighting. Nosy Be, Madagascar. Photograph taken on location for BBC Wild Madagascar Series, January 2010
Harlequin duck (Histrionicus histrionicus) male on a large boulder in the fast flowing and turbulent waters of the Laxa River, Iceland, May. Bookplate from Danny Greens The Long Journey North
Glass Frog (Cochranella mache) portrait, Ecuador, Endangered species
Red-bellied Lemur (Eulemur rubriventer) pair, Ialasatra, Madagascar
Male Coyote (Canis latrans) walking through deep snow. Hayden Valley, Yellowstone National Park, Wyoming, USA, February
Male Corncrake (Crex crex) South Uist, Outer Hebrides, Scotland, UK, June
