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Sperm whales (Physeter macrocephalus) aerial shot showing much larger male on right, with female nursery group. Baja California, Mexico
Snow leopard (Panthera uncia) old male walking on a cliff ledge in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Snow leopard (Panthera uncia) old male at dusk looking from a cliff ledge in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Tapanuli Orangutan (Pongo tapanuliensis). Togus, adult flanged male. Batang Toru Forest, Sumatran Orangutan Conservation Project, North Sumatran Province, Indonesia
West Africa lion (Panthera leo) subadult male resting on burnt ground, Pendjari National Park, Benin
Tompot blenny (Parablennius gattorugine) male peers out of its den, in leg of Swanage Pier, Swanage, Dorset, UK, English Channel, August
Eurasian blackcaps (Sylvia atricapilla) caught on limed stick, Cyprus. Finalist in The Wildlife Photojournalist Award: Single Image Category of the Wildlife Photographer of the Year Awards (WPOY)
Male Indri (Indri indri) leaping through the rain forest canopy. Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Andean Cock of the rock (Rupicola peruviana) male during courtship, Andean Mountains, Merida, Venezuela
Oribi (Ourebia ourebi montana), Senkelle Swaynes Hartebeest Sanctuary, Ethiopia
Banded demoiselle (Calopteryx splendens) two males on grass, Netherlands
Red-backed shrike (Lanius collurio) adult male taking off, Lower Saxony, Germany, June. Sequence 2 of 3
Lion (Panthera leo) male sharpening his claws on a tree at night, Sabi Sand Game Reserve, South Africa
Black bear (Ursus americanus) male close up head portrait, Minnesota, USA, June
Two wild Mustang stallions fighting in White Mountain Herd Area, Wyoming, USA. August
Wild Mustang stallion chasing another away from his family in White Mountain Herd Area, Wyoming, USA. August
Red Grouse (Lagopus lagopus scoticus) male on rock in swirling spindrift of snow, Scotland, UK, February. Highly commended in the Habitat category of the BWPA
Head portrait of grey Andalusian stallion running with mane flowing in arena in Northern France, Europe
White rhinoceros (Ceratotherium simum) two males fighting, in evening light, Sabi Sand Game Reserve, South Africa
Arctic grayling (Thymallus arcticus) displaying his vibrant dorsal fin. North Park, Colorado, USA
Ruff (Philomachus pugnax) males fighting at lek, Varanger, Norway, May 2016
Grey seal (Halichoerus grypus) young male plays and gapes, showing his teeth. Lundy IslandGrey seal (Halichoerus grypus) young male plays and gapes, showing his teeth.Lundy Island, Devon, UK, Bristol Channel, August
Close up of Red deer stag (Cervus elaphus) with antlers covered in velvet in spring, Daun, Vulkaneifel district, Rhineland-Palatinate, Germany, June. Captive
Ruff (Philomachus pugnax) male displaying, Varanger, Norway, May 2016
Alpine ibex (Capra ibex) portrait of young male, Bernese Alps, Switzerland, July
Young Sperm whale (Physeter macrocephalus) with erect penis Dominica, Caribbean Sea, Atlantic Ocean. Vulnerable species
Group of male Pochard (Aythya ferina) on water with motion blur, Welney WWT Reserve, Norfolk, England, UK, January
Silkmoth (Saturnia pyretorum) male, Miaoli, Taiwan
Male Common blue butterfly (Polyommatus icarus) Sutcliffe Park Nature Reserve, London. Eltham, London, UK. August
Male Common copper (Lycaena salustius) resting with wings open, Cape Kidnappers, Hawkes Bay, New Zealand, November
Emperor Moth (Saturnia pavoniella) male, on tree bark, Orvieto, Umbria, Italy, April
Golden eagle (Aquila chrysaetos) male head portrait beak open calling, Kalvtrask, Vasterbotten, Sweden. December
Close up Portrait of a Golden Eagle bird (Aquila chrysaetos) adult male, Kalvtrask, Vasterbotten, Sweden. Medium repro only
Long-tailed duck (Clangula hyemalis) Batsfjord village harbour, Varanger Peninsula, Norway, March
King eider duck (Somateria spectabilis) male, Batsfjord village harbour, Varanger Peninsula, Norway. March
American elk (Cervus elaphus canadensis) stag sticking tongue out, Yellowstone National Park, Wyoming, USA
Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines
Cow with cowbell silhouetted at sunrise, with mist over the Eastern Rhodope Mountains, Madzharovo, Bulgaria, November 2014
Southern elephant seal (Mirounga leonina), male at sunrise with King penguin (Aptenodytes patagonicus) colony in background. St Andrews Bay, South Georgia. October 2017
African wild dog (Lycaon pictus), male and female play fighting. Mana Pools National Park, Zimbabwe
Male Red-backed shrike (Lanius collurio), Cantabria, Spain, August
European stonechat (Saxicola rubicola) male perched with mountainous habitat behind, Sierra de Grazalema Natural Park, southern Spain, February
Mandarin duck (Aix galericulata) male swimming on water in the Beijing area, China, May
Lion (Panthera leo) close up portrait of male, captive occurs in Africa
African lion (Panthera leo) male lying in the middle of the Busanga Plains, Kafue National Park, Zambia. Vulnerable
Lion (Panthera leo) male resting, Masai Mara Game Reserve, Kenya, November
Black rhino (Diceros bicornis), male on the plains, Masai-Mara Game Reserve, Kenya
Jaguar (Panthera onca palustris) male resting on the banks of the Tres Irmaos River (Three Brothers River), a tributary of the Cuiaba River