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Four Hereford bullocks looking over a fence with Cow Parsley (Anthriscus sylvestris) Worcestershire, England, May
Moose (Alces alces) two bulls sparring in willow flat grasslands, Grand Teton National Park, Wyoming, USA. October
Banded demoiselle (Calopteryx splendens) male silhouetted on a fern, Lower Tamar Lakes, Cornwall, UK. July
Pair of Mute swans (Cygnus olor) at sunrise on misty lake. Poland, March
Fallow deer (Dama Dama) stags in velvet standing in a shady woodland spot, Knepp Estate, Sussex, UK, June
Male Common Redpoll (Acanthis flammea) perched on snow-covered dead stem of Evening Primrose (Oenothera biennis), from which it will eat seeds, New York, USA
Male Pine grosbeak (Pinicola enucleator) perched on branch among Crabapple (Malus sp) fruits, near Scenectady, New York, USA
Pied flycatcher (Ficedula hypoleuca), male, perched on a tree branch, Dumfries and Galloway, Scotland, UK. May
Male Jacksons chameleon (Chamaeleo jacksonii) on tree branch, East Africa
Scaly breasted lorikeet (Trichoglossus chlorolepidotus) pair allopreening in branch of tree, Toowoomba, Queensland, Australia
Eastern spinebill (Acanthorhynchus tenuirostris) male visiting Grevillea flower, small honeyeater found in forests, woodlands and gardens in Eastern Australia, Toowoomba, Queensland, Australia
Alpine ibex (Capra ibex) adult males fighting on a steep mountain side, Valsavarenche, Gran Paradiso National Park, Aosta Valley, Italy, May
Pine grosbeak (Pinicola enucleator) male feeding on Rowan (Sorbus sp) berries. Muurame, Central Finland. November
Wild boar (Sus scrofa) big male foraging in autumn forest during the hunting season in the Ardennes, Belgium. November
Ring-necked pheasant (Phasianus colchicus) male walking through leaflitter in autumnal colours in woodland on shooting estate, southern England, UK. November
Orange tip butterfly (Anthocharis cardamines) male resting in dappled light created by dew droplets. The Netherlands. April
Roe deer (Capreolus capreolus) feeding in field with flowering Poppies (Papaver rhoeas) and Cornflower (Centaurea cyanus), De Inslag, Brasschaat, Belgium, June
Himalayan ibex (Capra sibirica hemalayanus) males, resting and grazing. They live at elevations of 3800m and higher, western Himalaya mountains, Kibber Wildlife Sanctuary, India. April
Red deer (Cervus elaphus) stags sparring in the frost. Richmond Park, London, England, UK. December
Golden damsel (Amblyglyphidodon aureus) male aggressively defends his territory on a coral reef. Piaynemo Island, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Southern elephant seal (Mirounga leonina) male, rests and yawns on the snow. Right Whale Bay, South Georgia Island
Southern elephant seal (Mirounga leonina) male rests in a shallow lagoon. St Andrews Bay, South Georgia Island
Southern elephant seals (Mirounga leonina) male fighting for breeding rights. St Andrews Bay, South Georgia Island
Blyths paradise flycatcher (Terpsiphone affinis) male calling, Sabah, Malaysian Borneo
Proboscis monkey (Nasalis larvatus) male leaps between trees. Sabah, Malaysian Borneo
Pale-striped ground gecko (Lucasium immaculatum) male, from a desert sand plain near Mount Isa, North West Queensland, Australia. Controlled conditions
Kingfisher (Alcedo atthis) male perched on branch, Lorraine, France, July
RF - Long-tailed duck or Old-squaw (Clangula hyemalis) male, Varanger Peninsula, Norway, Scandinavia (This image may be licensed either as rights managed or royalty free.)
Magnificent hummingbird (Eugenes fulgens) male, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Red deer (Cervus elaphus) stag standing in bracken. London, UK. September
Northern spadefoot (Notaden melanoscaphus) adult male with vocal sac inflated, calling, Howard River sand flats, Northern Territory, Australia, January
Bilingual frog (Crinia bilingua) adult male calling, Adelaide River, Northern Territory, Australia, January
Long-tailed duck (Clangula hyemalis) male, Batsfjord, Norway, March
Kingfisher (Alcedo athis) perched on bull rush, Ubrique, Andalusia, Southern Spain, November
Southern chamois (Rupicapra pyrenaica) male, Picos de Europa, Spain, August
Portrait of a male Emerald glass frog (Espadarana prosoblepon) Ecuador
Stag beetle (Lucanus cervus) males fighting at dusk in oak wood, the Netherlands, June
Mandarin duck (Aix galericulata) male swimming with autumn colours reflected in the water, London, UK, November
Mandarin duck (Aix galericulata) male swimming, London, UK, November
Mallard (Anas platyrhynchos) male swimming with autumn colours reflected in the water, London, UK, November
Dingo (Canis lupus dingo) male, called Snapple, during a winter sunrise at the Dingo Dicovery and Research Centre. Snapple is an ambassador dingo for this often persecuted animal
African elephant (Loxodonta africana), male rubbing his trunk on termite hill, Samburu Game Reserve, Kenya
Gorilla (Gorilla beringei beringei) sub adult male feeding on vegetation, close up. Volcanoes National Park, Rwanda, Africa
Pine warbler (Dendroica pinus) male perched in blossoming Peach (Prunus persica) tree. Hill Country, Texas, USA
Blackpoll warbler (Setophaga striata) male perched on Mexican olive (Cordia boissieri). South Padre Island, Texas, USA
Ostrich (Struthio camelus) male preening, looking headless, Samburu Reserve, Kenya, Africa
Wild Alpine newt (Ichthyosaura alpestris) male, Lago Nero, Pistoia, Tuscany, Italy
Robin (Erithacus rubecula) male singing, perched on branch. Joutsa, Finland. April