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RF - Chalkhill blue butterfly (Lysandra coridon) male resting on Small scabious (Scabiosa columbaria). Somerset, England, UK. July 2019
RF - Red deer (Cervus elaphus) stag standing amongst Scots pine (Pinus sylvestris) tree trunks in forest. Cairngorms National Park, Scotland, UK. January
Swinhoes pheasant (Lophura swinhoii) male, endemic species, Yunlinshan, Taiwan
Black Woodpecker (Dryocopus martius), male feeding chicks, Bavaria, Germany
Masters snake (Drysdalia mastersii) male coiled on bush in mallee / heathland. Endangered local population due to repeated fires destroying habitat. Near Pinnaroo, Victoria, Australia
Nubian ibex (Capra nubiana), male standing in dry environment, Negev desert, Israel, April
Papuan hornbill (Rhyticeros plicatus) male perched in tree. Papua New Guinea
Sudells burrowing frog ( Neobatrachus sudellae ) male, A locally rare species from the Merri Creek corridor in northern metropolitan Melbourne, Victoria, Australia. Controlled conditions
Ring-necked pheasant (Phasianus colchicus) cock in wild flower meadow. Suffolk, UK. May
Warty / Clown frogfish (Antennarius maculatus) male portrait, Anilao, Batangas, Luzon, Philippines, March
Two wild konik horse stallions fighting in front of herd, Oostvaardersplassen Nature Reserve, Flevoland, Netherlands
Golden-browed chlorophonia (Chlorophonia callophrys), two males perched on moss covered branch. Savegre Valley. Costa Rica
Shining honeycreeper (Cyanerpes lucidus) male and Red-legged honeycreeper (Cyanerpes cyaneus) perched on branch with epiphyte. Boca Tapada, Costa Rica
Black-bellied bustard (Lissotis melanogaster) male calling at sunrise. Serengeti National Park, Tanzania
Andean Cock-of-the-rock male (Rupicola peruvianus) at lek in cloud forest canopy. Manu Biosphere Reserve, Amazonia, Peru. November
Stag beetle (Lucanus cervus) silhouetted at sunset. The Netherlands. August
Red Deer (Cervus elaphus) Stag in the snow. Scotland, March
Barred warbler (Sylvia nisoria) male perched in tree, surrounded by yellow flowers. Pargas, Aboland, Finland. May
Black woodpecker (Dryocopus martius) male on tree trunk, portrait. Kuusamo, Northern Ostrobothnia, Finland. February
Yellowhead jawfish (Opistognathus aurifrons) male rises up from its burrow and aerates the eggs he is mouthbrooding. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Sulawesi black macaques (Macaca nigra) males foraging on exposed beach at low tide. Tangkoko National Park, Sulawesi, Indonesia
RF - Elderly male snow leopard (Panthera uncia) stalking Siberian ibex (Capra sp. )RF - Elderly male snow leopard (Panthera uncia) stalking Siberian ibex (Capra sp.). Spiti Valley, Himachal Pradesh, northern India
Lusitano horses, five young stallions playing between trees in pasture, silhouetted at dawn. Portugal. April
Red fox (Vulpes vulpes) young male cub near entrance to earth in woodland, Switzerland
Red deer stag (Cervus elaphus) and a Holm oak tree (Quercus ilex) silhouetted Parque Natural Sierra de Andujar, Andalucia, Spain. January
Young Roe deer (Capreolus capreolus) buck with developing horns in velvet approaching a garden pond, Wiltshire garden, UK, February
Portrait of male silverback Mountain gorilla (Gorilla beringei beringei) feeding. Bwindi Impenetrable Forest National Park, Uganda, Africa
Chimpanzee (Pan troglodytes schweinfurthii) male vocalising in tree, Kibale National Park, Uganda, Africa
Mountain gorilla (Gorilla berengei), aggressive young male from Titus Group, Visoke, Volcanoes National Park, Rwanda, August
Galapagos vermilion flycatcher (Pyrocephalus nanus) male perched with fledglings, Alcedo Volcano, Isabela Island, Galapagos
Galapagos sea lions (Zalophus wollebaeki) sub adult males playing with starfish, Mosquera Islet, Galapagos
Santa Cruz lava lizard (Microlophus indefatigabilis), Santa Fe Island, Galapagos
Scorpion fly (Panorpa sp. ) male, basking in foliage. Peak District National ParkScorpion fly (Panorpa sp.) male, basking in foliage. Peak District National Park, Derbyshire, UK. May
Portrait of male 12 year old domestic cat, ragdoll breed, bicolour, named Magellan, in grass looking at camera. Brighton, Victoria, Australia
Yellow Warbler (Setophaga petechia, formerly Dendroica petechia), male in breeding plumage, Ithaca, New York, USA. May 2020
Wood duck (Aix sponsa) male in breeding plumage. Acadia National Park, Maine, USA. April
Resplendent quetzal (Pharomachrus mocinno), male and female, Talamanca mountains, Costa Rica
Anolis lizard (Anolis sp. ) male displaying in Corcovado National Park, Costa RicaAnolis lizard (Anolis sp.) male displaying in Corcovado National Park, Costa Rica
RF - Sand lizard (Lacerta agilis) male on sand dune, the Netherlands. May. (This image may be licensed either as rights managed or royalty free.)
Sand lizard (Lacerta agilis) male, the Netherlands
Blue sheep (Ovis nahoor) male. Valley of the Cats, Angsai Nature Reserve, Sanjiangyuan National Nature Reserve, Tibetan Plateau, Qinghai, China. October
Three-toed woodpecker (Picoides tridactylus), male next to nest hole in tree, Finland, June
Cuckoo wrasse (Labrus mixtus) male in front of Red sea fan, (Paramuricea clavata), Stupiste In dive site, Vis Island, Croatia, Adriatic Sea, Mediterranean
Leucistic white lions (Panthera leo krugeri) male and juveniles, Captive
Crested dragon (Ctenophorus cristatus) male basking on a fallen log in chenopod shrubland habitat near Balladonia, Nullarbor Plain, Western Australia
Four Pygmy seahorses (Hippocampus bargibanti) on a Seafan / Gorgonian (Muricella sp. ) including one or several pregnant males. New Caledonia, Pacific Ocean
Vagrant darter (Sympetrum vulgatum) male resting. Brasschaat, Belgium. August
African elephant (Loxodonta africa) male in threat display to rival. Floor of the Ngorongoro Crater, Ngorongoro Conservation Area, Tanzania