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Jumping spider (Euophrys frontalis) male amongst flower petals, Peak District National Park, Derbyshire, UK, June
Male Indian peafowl (Pavo cristatus) displaying, captive
Male Whinchat (Saxicola rubetra) perched on Oil seed rape (Brassica napus) in a field, Vosges, France, April
A rare Sorraia stallion cantering, Coudelaria Nacional (National Stud Farm), in Alter do Chao, District of Portalegre, Alentejo, Portugal
Jaguar (Panthera onca palustris) male wading through water at the shore of the Piquiri River. The Pantanal wetlands of Mato Grosso, Mato Grosso State, Brazil, October
African elephants (Loxodonta africana) bulls crowd into an emptying waterhole to drink, Savuti, Botswana. Taken on location for BBC Planet Earth series, 2005
Pheasant (Phasianus colchicus) male amongst Bluebell flowers, UK, April
Two male Mute swans (Cygnus olor) displaying on water in territorial dispute, UK
A male Leafy Seadragon (Phycodurus eques) swimming. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Chestnut Paso peruvian stallion running, California, USA
Hoolock Gibbon (Hylobates hoolock) male hanging from a branch. Assam, India, February
Wild stallion (Equus caballus) lying down in the sagebrush and yawning. The foothills south of Reno, Nevada, USA, May
Red Deer (Cervus elaphus) male standing in snow. The Netherlands, January
Five Mallard (Anas platyrhynchos) walking in a line across snow. Three males and two females. The Netherlands, January
Red deer (Cervus elaphus) stag amongst winter vegetation, Salburua Park, Alava, Spain, November
African lion (Panthero leo) male giving flehmen grimace, Tanzania, East Africa
Feral domestic horses (Equus caballus) wild herd stallion (grey) fighting a bachelor chestnut stallion, Blue Swallow Reserve, near Kaapsehoop, Mpumalanga province / Eastern Transvaal, South Africa
Polar bear (Ursus maritimus) portrait of curious young male, covered in snow from rolling around / playing. On a barrier island during autumn freeze up, Bernard Spit
Harlequin duck (Histrionicus histrionicus) portrait of male stood motionless on exposed rock, in river, Iceland, May
Ptarmigan (Lagopus mutus) male in flight over snow covered Cairngorm Mountains, Scotland, February. (non-ex)
Old male African lion (Panthera leo) portrait, Sabi Sand Game Reserve, South Africa, June
Giraffe necked weevil {Trachelophorus giraffa} male on leaf of host plant (Dichaetanthera cordifolia) tropical rainforest, Mantadia NP, Madagascar
Polar bear (Ursus maritimus) head close-up portrait of an adult male, with snowflakes on fur, Barter Island, 1002 area of the Arctic National Wildlife Refuge, Alaska
Dalmatian, head portrait, male aged 4 years
Adult male Bornean orangutan (Pongo pygmaeus) in rainforest canopy, Gunung Palung National Park, Borneo, West Kalimantan, Indonesia
HeaPair of male Boxers fawn coloured with natural ears, sitting by lake in Autumn, Illinois, USA
A Sika deer stag rests among a band of wild Misaki-uma horses (Equus ferus caballus) in the Cape Toi Reserve, Miyazaki Prefecture, Kyushu Island, Japan
A wild Misaki-uma breeding stallion (right) brings back one of his mares to his band while another breeding stallion (left) charges him away, in the Cape Toi Reserve, Miyazaki Prefecture
Male American robin (Turdus migratorius) puffed up to keep warm, and perched on Juniper branch, Kentucky, USA
Head portrait of male Mandrill (Mandrillus sphinx) Captive. Apenheul zoo; the Netherlands
African lion (Panthera leo) male and female at sunrise, Masai Mara National Reserve, Kenya, January
Male African lion (Panthera leo) lying in grass at sunrise, Masai Mara National Reserve, Kenya, January
Two Elk (Cervus canadensis) bulls at sunset, Jasper National Park, Rocky Mountains, Alberta, Canada, September
Two male Thomsons gazelles (Eudorcas thomsonii) fighting, Masai-Mara game reserve, Kenya
Roe deer (Capreolus capreolus) male portrait, , Vosges, France, May
Male Andean cock of the rock (Rupicola peruvianus) perched on branch, Mindo, Pichincha, Ecuador
