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Vogelkop bowerbird (Amblyornis inornatus) male, at his bower surrounded by orange leaves and petals and other objects he has collected, Arfak Mountains, Vogelkop Peninsula, West Papua
Wandering albatross (Diomedea exulans) male, standing with wings spread in courtship display, Prion Island, South Georgia
Magnificent riflebird (Ptiloris magnificus alberti) male, courtship display, Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
Mountain gorilla (Gorilla gorilla beringei) silverback, head portrait, Bwindi Impenetrable Forest National Park, Uganda, Africa. Critically endangered
Male Pygmy killer whale (Feresa attenuata) swimming in open ocean, Hawaii, Pacific Ocean
Male Highland Great diving beetle (Dytiscus lapponicus) on surface of water, Darnaway Estate, Highlands, Scotland, UK. May. Focus stacked. Captive
Kerry bog pony, stallion, a rare breed, running, County Kerry, Republic of Ireland. April
Falabella miniature horse, stallion, walking over grassland, County Kerry, Republic of Ireland. April
Anadolu stallion standing alert, portrait, Sirince mountains, Turkey
Kerry bog pony, gelding, a rare breed, head portrait, County Kerry, Republic of Ireland. April
Pair of Lava gulls (Leucophaeus fuliginosus) during courtship, Genovesa Island, Galapagos Islands, Ecuador
Male Banded demoiselle (Calopteryx splendens), abstract study of wings and body, Tamar Lakes, Cornwall, UK. June
Male Australian magpie (Gymnorhina tibicen) standing on stump with beak full of moths, backlit. Canterbury, South Island, New Zealand
Komodo dragon (Varanus komodoensis) male, foraging in mangove / inter-tidal zone at low tide, Komodo Island, Komodo National Park, Indonesia. Endangered
Two Komodo dragons (Varanus komodoensis) males, fighting on a remote beach, Komodo Island, Komodo National Park, Indonesia. Endangered
Young male Lion (Panthera leo) resting in tree during middle of the day to escape the heat. Acacia woodland near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border, Tanzania
Young male Lion and lioness (Panthera leo) resting in tree during middle of the day to escape the heat. Acacia woodland near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border
Northern parula (Parula americana) male singing from flowering crabapple (Malus sp)Northern parula (Parula americana) male singing from flowering crabapple (Malus sp.) tree in spring, near Salamanca, New York, USA, May 2022
Redstart (Phoenicurus phoenicurus) juvenile male, in flight, chasing an insect, Parainen Uto, Finland. September
King eider (Somateria spectabilis) male, on water, portrait, Batsfjord, Norway. February
Hen harrier (Circus cyaneus) juvenile male, in flight, Parainen Uto, Finland. September
Male Cheetah (Acinonyx jubatus) stretching and yawning on a fallen tree trunk, Okavango Delta, Botswana, Africa
Two male Common / Plains zebras (Equus quagga) fighting, Okavango Delta, Botswana, Africa
Male Gold-specs jawfish (Opistognathus randalli), with mouth brooding eggs, the eyes can clearly be seen in the eggs indicating they will soon hatch, Mabul Island, Sabah, Malaysia, Celebes Sea
Male Eastern bluebird (Sialia sialis) feeding on snow-covered Holly (Ilex sp) berries, near Ithaca, New York, USA. January
Male Orange tip butterfly (Anthocharis cardamines) resting on Wild garlic (Allium ursinum) flower, Peak District National Park, Derbyshire, UK. April
Male Jumping spider (Euophrys frontalis) close up, Derbyshire, UK. May. Focus stacked image
Male Giant kingfisher (Megaceryle maxima) diving, Allahein River, The Gambia
Male Beautiful sunbird (Cinnyris Pulchellus) courtship display, Allahein River, The Gambia
Three male Ostriches (Struthio camelus) running and flapping wings in aggressive display, Kgalagadi Transfrontier Park, South Africa
Male Jackson's chameleon (Chamaeleo jacksoni) reaching out as it moves between leaves, Maui, Hawaii
RF - Male Blackhead blenny (Emblemariopsis bahamensis) peering out from a hole in a Brain coral (Scleractinia), Grand Cayman, Cayman Islands, Caribbean Sea
RF - Male Dugong (Dugong dugon) swimming beneath the surface with a young Golden trevally (Gnathanodon speciosus) alongside, Marsa Alam, Egypt, Red Sea
RF - Male Great white shark (Carcharodon carcharias) swimming beneath the surface, Guadalupe Island, Baja California, Mexico, Pacific Ocean
Male Sailfin blenny (Emblemaria pandionis) displaying large dorsal fin, portrait, Grand Cayman, Cayman Islands, Caribbean Sea
Pair of Peppermint basslets (Liopropoma rubre) spawning at sunset, with eggs floating alongside the female, Grand Cayman, Cayman Islands, Caribbean Sea
Male Emperor angelfish (Pomacanthus imperator) swimming over garden of Fire corals (Millepora sp)Male Emperor angelfish (Pomacanthus imperator) swimming over garden of Fire corals (Millepora sp.) in morning sunshine, Ras Mohammed National Park, Sinai, Egypt, Red Sea
Two male Black grouse (Tetrao tetrix) fighting for territory on lek, Cairngorms National Park, Scotland, UK. May
Black-faced sheep (Ovis domesticus) ram, head portrait, Isle of Islay, Hebrides, Scotland, UK. April
Two male Polar bears (Ursus maritimus) standing on hind legs, sparring, Churchill, Canada. November
Two Red deer (Cervus elaphus) stags fighting, Bradgate Park, Leicestershire, UK. October
Soft focus close up of male Banded demoiselle damselfly (Calopteryx splendens) body and wings, roosting among grasses. Cornwall, UK. July
Male Kingfisher (Alcedo atthis) sitting on eggs for his brooding period in an artificial nest, Italy
Male Kingfisher (Alcedo atthis) feeding chicks, aged 9 days, with a large fish in artificial nest, Italy
Camargue bull (Bos taurus) running through marshland, Camargue, France. October
Red deer (Cervus elaphus) stag jumping over a stock fence along roadside, Cairngorms National Park, Scotland, UK. February
Red deer (Cervus elaphus) stag walking through falling snow, Cairngorms National Park, Scotland, UK. January
Male Black grouse (Tetrao tetrix) performing courtship display at lek, Cairngorms National Park, Scotland, UK. April