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Glass Frog (Cochranella mache) portrait, Ecuador, Endangered species
Red-bellied Lemur (Eulemur rubriventer) pair, Ialasatra, Madagascar
Male Coyote (Canis latrans) walking through deep snow. Hayden Valley, Yellowstone National Park, Wyoming, USA, February
Male Corncrake (Crex crex) South Uist, Outer Hebrides, Scotland, UK, June
Bengal Tigers (Panthera tigris) sub-adults, approximately 17-19 months old, playfighting. Endangered. Bandhavgarh National Park, India. Non-ex. Sequence 1 of 2
Lion (Panthera Leo) male running, South Africa taken on location for Pride tv series
Mallard duck (Anas platyrhynchos) drake swimming through reflection of house, Bude canal, Cornwall, UK. April 2015
Speckled Wood (Pararge aegeria) male flying in habitat, Finland, April
Herd of Camargue bulls, Camargue, France, May
Bearded tit (Panurus biarmicus) male in summer plumage. Hungary, May
Wild male Jaguar (Panthera onca palustris) stalking along the bank of the Cuiaba River in late afternoon sun light. Northern Pantanal, Brazil
Golden Retriever, male in straw
Japanese Macaque (Macaca fuscata) young one, appearing quite relaxed in thermal pool, even falling asleep, in Jigokudani, Japan, February
Male Long-tailed duck (Clangula hyemalis), Batsfjord, Norway, March
Group of male Common eiders (Somateria mollissima) in flight, Iceland, June
Male African lion (Panthera leo) at night, Kalahari Desert, Botswana
Bighorn sheep (Ovis canadensis) male ram walking in the snow, Yellowstone National Park, USA, February
Black bear (Ursus americanus) male resting in grass meadow, Minnesota, USA, June
Moose Bull (Alces alces) walking in forest clearing, Denali National Park, USA, September
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Hippo (Hippopotamus amphibius) male after a a mud bath, Masai-Mara Game Reserve, Kenya. Vulnerable species
Large male Reindeer (Rangifer tarandus) standing in shallow waters of the Kronotsky lagoon, away from biting insects. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Male Little bustard (Tetrax tetrax) displaying, Catalonia, Spain, May
Lion (Panthera leo) males resting, Masai-Mara Game Reserve, Kenya. Vulnerable species
Lion (Panthera leo) male using flehmen response to smell, Masai-Mara Game Reserve, Kenya. Vulnerable species
Lion (Panthera leo) male, Masai-Mara Game Reserve, Kenya. Vulnerable species
Male Choco trogon (Trogon comptus) perched on branch, Mashpi, Pichincha, Ecuador
Male Masked trogon (Trogon personatus) portrait, Mindo, Pichincha, Ecuador
Male Collared trogon (Trogon collaris) perched on twig, Sumaco National Park, Napo, Ecuador
Tapichalaca torrenteer frog(Hyloscirtus tapichalaca) male on a bromeliad, Tapichalaca Reserve, Loja, Ecuador
Andean Cock-of-the-Rock (Rupicola peruvianus) Mindo, Pichincha, Ecuador
RF - Snow leopard (Panthera uncia) male moving over rocky, snow-covered slopes. Ulley Valley, Himalayas, Ladakh, India
Great horned owl (Bubo virginianus) male roosting in an abandoned barn. Idaho, USA. February
Red Deer Stag (Cervus elaphus) amongst Autumn vegetation, Bradgate Park, Leicestershire, UK, October 2015
Ring-necked pheasant (Phasianus colchicus) male standing among bluebells during spring in woodland on shooting estate, southern England, UK, April
Pair of Giant frogfish (Antennarius commersoni), male on the left, female above on right on sponges. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
RF- Northern Goshawk (Accipiter gentilis) male close-up, Scotland
Portrait of male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water above coral reef, with bokeh effect in water from the sunset. The Alternatives, Sinai, Egypt
RF - Shallow depth of field portrait of male Scalefin anthia (Pseudanthias squamipinnis) swimming in open water above coral reef photographed against sunset. The Alternatives, Sinai, Egypt
Tiger cardinalfish (Cheilodipterus macrodon) male mouthbrooding eggs in his mouth. Gubal Barge, Gubal Island, Egypt. Strait of Gubal, Gulf of Suez, Red Sea
A batch of Leafy Seadragon (Phycodurus eques) eggs on the tail of a male. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
Andean cock of the rock (Rupicola peruviana) male in courtship ritual, Andean Mountains, Merida, Venezuela
Stellers eider duck (Polysticta stelleri) Batsfjord village harbour with red reflections on the water, Varanger Peninsula, Norway, March
Pine Grosbeak (Pinicola enucleator) male feeding on berries with traffic light behind, Oulu, Finland, December
White-tailed Deer (Odocoileus virginianus) in front of forest, with reflection of trees in water, Acadia National Park, Maine, USA, October
Pine grosbeak (Pinicola enucleator) young male feeding on rowan berries covered in snow, Liminka, Finland. November
Banded jawfish (Opistognathus macrognathus) male incubating eggs in mouth, which are at a late stage of development and will soon be ready for release
Jumping spider (Euophrys frontalis) male amongst flower petals, Peak District National Park, Derbyshire, UK, June
