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Red deer stag (Cervus elaphus) with female during rut, Exmoor National Park, Somerset / Devon, England. November
Two male Black grouse (Lyrurus tetrix) fighting and sparing over territory at a snow covered lek, Vasterbotten, Sweden. April
Common pheasant (Phasianus colchicus) male flapping wings and calling, Pusztaszer reserve, Hungary. May
Two young male African forest elephants (Loxodonta cyclotis) fighting over small pool of water in Dzanga-Sangha National Park, Central African Republic
Male Moose (Alces alces) with antlers, portrait, Finland. September
Male Mountain apollo butterfly (Parnassius apollo) nectaring on wildflower, Finland. July
Alpha male Grey wolf (Canis lupus) biting angry young wolf to assert dominance, Finland. July
Young male Collared flycatcher (Ficedula albicollis) in between budding branches, Finland. May
Close up of male Western capercaillie (Tetrao urogallus) during lek mating, Finland. March
Male Swallowtail butterfly (Papilio machaon) eating from Wood cranesbill (Geranium sylvaticum), Finland. June
Male Yellowface pike-blenny (Chaenopsis limbaughi), living in abandoned tubeworm holes in sand, flaring its dorsal fin in threat display, Dominica, Eastern Caribbean
Male Giant kingfisher (Megaceryle maxima) perched on branch, Allahein river, The Gambia
Male Spanish moon moth (Graellsia isabellae) close up of forewing eyespot, Switzerland
Mandarin duck (Aix galericulata) drake swimming on pond, backlit by rising sun, Richmond Park, London, UK. February
Two male Alpine Ibex (Capra ibex) fighting on a mountain slope at dawn, Varaita Valley, Alps, Italy, July
Male Blackbird (Turdus merula) feeding on berries in Hawthorn (Crataegus sp. ) hedgerow, Norfolk, UK. NovemberMale Blackbird (Turdus merula) feeding on berries in Hawthorn (Crataegus sp.) hedgerow, Norfolk, UK. November
Three Lined seahorses (Hippocampus erectus), two male and one female, feeding on plankton in an alkaline saltwater pond, The Bahamas, Caribbean
Male Bearded tit (Panurus biarmicus) perched on a reed stem, Leighton Moss Nature Reserve, Lancashire, UK. September
Male Ring ouzel (Turdus torquatus) standing on a garden lawn, Norfolk, UK. April
Red deer (Cervus elaphus) stag amongst ferns, Bradgate Park, Leicestershire, UK. October
Matriarch Timber wolf (Canis lupus lycaon) sitting next to alpha male standing in snow. Hudson Bay, Manitoba, Canada
Orchard oriole (Icterus spurius) male feeding on Lemon bottlebrush (Callistemon pallidus) flowers. South Padre Island, Texas, USA
Ladder-backed woodpecker (Picoides scalaris) male perched on blossoming Mexican plum (Prunus mexicana). Hill Country, Texas, USA
RF - Banded demoiselle damselfly (Calopteryx splendens) male roosting on reed, silhouetted against sun reflected in the water. Cornwall, England, UK. May
Red deer (Cervus elaphus) stag bellowing amongst bracken, Bushy Park, London, England, UK, October
Lions (Panthera leo) - two brothers patrolling territorial boundary at border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Male Resplendent Quetzal (Pharomachrus mocinno) in cloud forest. Los Quetzales National Park, Savegre River Valley, Talamanca Range, Costa Rica, Central America
Male Violet Sabrewing (Campylopterus hemileucurus) hovering / in flight sequence. Montane forest, Bosque de Paz, Caribbean slope, Costa Rica, Central America
Male Bare-faced Curassow (Crax fasciolata) near Araras Lodge, Mato Grosso, Northern Pantanal, Brazil, South America
RF- Adult male Serval (Felis serval) stalking prey in long grass. Serengeti National Park, Tanzania
Male Blackbird (Turdus merula) feeding on berries in winter, Bavaria, Germany, Europe. January
Luminous, glowing light tracks from male Fireflies (Lamprohiza splendidula) in the forest at dusk, Bavaria, Germany, Europe. July
Male Lesser spotted woodpecker (Dryobates minor) perching on tree trunk, Germany, Europe. May
Bull Moose (Alces alces) drinking from mountain stream, Grand Teton National Park, Wyoming, USA. October
Dominant male Chimpanzee (Pan troglodytes troglodytes) walking in the mangrove, with female carrying her infant, Conkouati-Douli National Park, Republic of Congo, Africa
Male Dendtric jawfish (Opistognathus dendriticus) with incubating eggs in his mouth, Dampier Strait, Raja Ampat, Indonesia, Pacific Ocean
Female Yellow-footed rock wallaby (Petrogale xanthopus) fending off advances from male (right), Idalia National Park, Queensland, Australia
Male Great frigatebird (Fregata minor) in breeding condition with semi-inflated throat pouch, Christmas Island, Australia
Dominant male Chimpanzee (Pan troglodytes troglodytes) in the mangrove, with female and infant vocalising, Conkouati-Douli National Park, Republic of Congo, Africa
Male Saiga antelope (Saiga tatarica) rutting in winter, The Black Lands National Park, Kalmykia, Russia
Two male Darjeeling bush frogs (Raorchestes Annandalii) fighting over territory, Kalimpong, West Bengal, India
Male Santiago lava-lizard (Microlophus jacobi), among Galapagos carpetweed (Sesuvium edmonstoni), Bainbridge Islets, Galapagos, South America
Male Bulwers pheasant / Bulwers wattled pheasant (Lophura bulweri) endemic to the forests of Borneo, Pairi Daiza, Belgium. Captive, occurs in Borneo
Wild Welsh pony colt greeting his father, Carneddau Mountains, Snowdonia, Wales, UK. June
Mexican fiddler crab (Leptuca crenulata) small male displaying with larger individual behind, Morua Estuary, near Rocky Point (Puerto Penasco), Gulf of California, Mexico, June
Male Bleekers parrotfish (Chlorurus bleekeri) races across a coral reef as it displays to females.Yanggefo Island, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Male Lumpsucker (Cyclopterus lumpus) swimming through Kelp forest (Laminaria hyperborea), Loch Carron, Highlands, Scotland, UK
Male Jaguar (Panthera onca) walking through rainforest, Belize, Central America, 2017. Captive. Filmed for the BBC series Big Cats
