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African elephant (Loxodonta africa) male in threat display to rival. Floor of the Ngorongoro Crater, Ngorongoro Conservation Area, Tanzania
RF - Male African lion (Panthera leo) scratching tree at Big Marsh, Nogorongoro Conservation Area / Serengeti National Park, Tanzania. March
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
Haswells frog (Paracrinia haswelli) male, Pambula Beach, New South Wales, Australia. Controlled conditions
Polar bear (Ursus maritimus) male approaching female in rocky area covered in snow. Svalbard, Norway, April 2019
Southern elephant seal (Mirounga leonina), two males, equally matched with mouths open in aggression. Out of focus King penguin (Aptenodytes patagonicus) colony in background
Red deer (Cervus elaphus) stag bellowing during rut in grassland, doe in background
Dalmatian wall lizard, (Podarcis melisellensis), fight between two males with distinct morphs, Croatia, April. Non-ex
Orange breasted sunbird (Anthobaphes violacea), male nectaring on Heather (Erica sp). Kirstenbosch Botanic Garden, Cape Town, South Africa. February
Narcissus fly (Merodon equestris) male nectaring on Rock rose (Cistus x hybridus) in garden, Surrey, England, UK
Carpenter bee (Xylocopa caffra) male robbing nectar from Ninepin heath (Erica mammosa), not pollinating flower. The Cape, South Africa. February
Pere Davids deer (Elaphurus davidianus) two stags boxing amongst herd of hinds. Captive bred in reserve after reintroduction from Woburn Abbey, England, UK
Great eggfly (Hypolimnas bolina) butterfly male nectaring on Rough-leaved hydrangea (Hydrangea aspera). Male is highly territorial chasing other insects away
Red-flanked bluetail (Tarsiger cyanurus) male front view, Jyvaskyla, Central Finland, May
Red deer (Cervus elaphus) stags in snowy Pine forest. Scotland, UK. March
Blackbird (Turdus merula) male in winter, Bavaria, Germany, December
Drill (Mandrillus leucophaeus) male portrait, captive
African lion (Panthera leo) young male, portrait, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa. A
Red deer (Cervus elaphus) stag roaring, silhouetted under trees at sunrise. Bushy Park, London, UK. September
Red Deer (Cervus elaphus) stag after a mud bath, Bradgate Park, Leicestershire, UK, October
Male Alpine ibex (Capar Ibex) resting, Massif des Cerces, Savoie France, October
Fallow deer (Dama dama) herd at dawn, Holkham, Norfolk, England, UK, October
RF- Northern Cardinal (Cardinalis cardinalis) immature male perched on Texas Red Oak (Quercus Texana). New Braunfels, San Antonio, Hill Country, Central Texas, USA
RF - Roe buck (Capreolus capreolus) stood in rough grassland in evening light, Scotland. September
Red deer (Cervus elaphus) stag bellowing, surrounded by hinds, Surrey, UK, September
Red deer, (Cervus elaphus), stag in rough grassland, Jura, Scotland, UK. April
Banded demoiselle (Calopteryx splendens) male on yellow lotus ( Nuphar lutea) Norfolk, England, UK, August
Red Deer (Cervus elaphus) male with new antler growth, Minsmere RSPB Reserve, Suffolk, England, UK, August
European pochard (Aythya ferina) drake male on a misty morning, Valkenhorst Nature Reserve, Valkenswaard, The Netherlands, May
Chimpanzee male (Pan troglodytes schweinfurthii) sleeping on the forest floor with feet up, Kibale National Park, Uganda. January
White-backed woodpecker (Dendrocopos leucotos) male on tree trunk. Oulu, Finland. January
Citrine wagtail (Motacilla citreola) male perched on Reed. Latvia. April
Hairy woodpecker (Picoides villosus) male in snowstorm, Freeville, New York, USA, February
Male Downy woodpecker (Picoides pubescens) excavating nest cavity, tossing wood chips out of nest hole in tree trunk, Lansing, New York, USA, May
Sperm whales (Physeter macrocephalus) aerial shot showing much larger male on right, with female nursery group. Baja California, Mexico
Snow leopard (Panthera uncia) old male walking on a cliff ledge in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Snow leopard (Panthera uncia) old male at dusk looking from a cliff ledge in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Tapanuli Orangutan (Pongo tapanuliensis). Togus, adult flanged male. Batang Toru Forest, Sumatran Orangutan Conservation Project, North Sumatran Province, Indonesia
West Africa lion (Panthera leo) subadult male resting on burnt ground, Pendjari National Park, Benin
Tompot blenny (Parablennius gattorugine) male peers out of its den, in leg of Swanage Pier, Swanage, Dorset, UK, English Channel, August
Eurasian blackcaps (Sylvia atricapilla) caught on limed stick, Cyprus. Finalist in The Wildlife Photojournalist Award: Single Image Category of the Wildlife Photographer of the Year Awards (WPOY)
Male Indri (Indri indri) leaping through the rain forest canopy. Andasibe-Mantadia National Park, eastern Madagascar. Endangered
Andean Cock of the rock (Rupicola peruviana) male during courtship, Andean Mountains, Merida, Venezuela
Oribi (Ourebia ourebi montana), Senkelle Swaynes Hartebeest Sanctuary, Ethiopia
Banded demoiselle (Calopteryx splendens) two males on grass, Netherlands
Red-backed shrike (Lanius collurio) adult male taking off, Lower Saxony, Germany, June. Sequence 2 of 3
Lion (Panthera leo) male sharpening his claws on a tree at night, Sabi Sand Game Reserve, South Africa
Black bear (Ursus americanus) male close up head portrait, Minnesota, USA, June
