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Pale-mandibled aracaris (Pteroglossus erythropygius) pair, at entrance to nest hole in tree, Choco rainforest, Los Bancos, Pichincha, Ecuador
Club-winged manakin (Machaeropterus deliciosus) male, courtship display at lek, Choco rainforest, Pichincha, Ecuador
Great spotted woodpecker (Dendrocopos major) male parent feeding juvenile. Germany. June
Brambling (Fringilla montifringilla) male, perched on a lichen covered branch. Andalusia, Spain. October
African elephant (Loxodonta africana), male, covered in red dust, under shade of tree. Tsavo East National Park, Kenya
RF - Orange tip butterfly (Anthocharis cardamines) male, resting on flower (Cruciferae sp)RF - Orange tip butterfly (Anthocharis cardamines) male, resting on flower (Cruciferae sp.), Peerdsbos, Brasschaat, Belgium. May. (This image may be licensed either as rights managed or royalty free.)
Male Siberian flying squirrel (Pteromys volans orii) attempting to enter nest with female inside during reproductive season but stopped mid-way then rejected. Hokkaido, Japan. March
Pair of Japanese dwarf flying squirrels (Pteromys volans orii) during the reproductive season, on branch nlooking down, Hokkaido, Japan. The female is on the left, male right
Male Korean seahorse (Hippocampus haema) giving birth. Kumamoto Prefecture, Kyushu, Japan
Humpback whale (Megaptera novaeangliae) rear view of males escort, with adult female and male calf in the background. Vava'u, Tonga, South Pacific
Killer whale / Orca (Orcinus orca) mature male, swimming underwater. Hamn, Senja, Norway, Atlantic Ocean
Humpback whale (Megaptera novaeangliae) male calf with his mother. Vava'u, Tonga. South Pacific
Australian sea lion (Neophoca cinerea) male with Striped cowfish (Aracana aurita) Carnac Island, Western Australia
Indri (Indri indri) male, leaping through the rain forest canopy, Andasibe-Mantadia National Park, eastern Madagascar. Critically Endangered
Fosa (Cryptoprocta ferox) pair, mating in forest canopy, Zombitse-Vohibasia National Park, central south western Madagascar
Crowned lemur (Eulemur coronatus), male, climbing over limestone karst rock formations (tsingy), Ankarana Special Reserve, Madagascar. Endangered
Pair of Kestrels (Falco tinnunculus) perched on tree stump in rain, UK. March
Kestrel (Falco tinnunculus) male, perched on fallen branch, UK. May
Alpine ibex (Capra ibex) male, with kid on mountain ridge in heavy winter snowfall, Valsavarenche, Gran Paradiso National Park, Italy. January
Black grouse (Lyrurus tetrix) male, jumping, part of a dominance display, at lek in early morning light, Kuusamo, Finland, April
Hairy dragonfly (Brachytron pratense) male, resting on plant stem, Brackagh Moss National Nature Reserve, County Armagh, Northern Ireland, UK. June
Jumping spider (Evarcha arcuata), male, in alert pose, ready to jump. Buckinghamshire, England. UK. August. Focus Stacked
Aggregation of Sperm whales (Physeter macrocephalus) several females swim after a few males, Dominica, Caribbean Sea, Atlantic Ocean. Photo taken under permit. February
Velvet swimming crab (Necora puber) male, in defensive posture, with wreck of the Rosalie in background, Weybourne, Norfolk, England, UK, North Sea
Pair of Spotted unicornfish (Naso brevirostris) courtship display above the reef, Ari Atoll, Maldives, Indian Ocean
Scalefin anthias (Pseudanthias squamipinnis) male, swimming close to surface, Sharm El Sheikh, Sinai, Egypt, Red Sea
Pair of Nudibranchs (Flabellina exoptata) just prior to mating, the mating apparatus of the one on left is visible, Bitung, North Sulawesi, Indonesia, Molucca Sea
Two Dartmoor ponies (Equus ferus caballus), endangered rare breed, colts greeting one another. Dartmoor National Park, Devon, England. June
Dartmoor pony (Equus ferus caballus), endangered rare breed, mare and colt standing alert. Dartmoor National Park, Devon, England. June
Asian elephant (Elephas maximus indicus) male, eye detail through foliage, Bardia National Park, Terai, Nepal
Red-backed toadlet (Pseudophryne coriacea) male, guarding eggs in nest under rotten log, Border Ranges, Queensland, Australia
Two Red deer (Cervus elaphus) stags, fighting during the rut, Cheshire, UK. July
Two Ruffs (Philomachus pugnax) male, competing at a lek, Vardo, Norway. May
Blackburnian warbler (Dendroica fusca) male in breeding plumage, on branch with newly-emerging leaves in spring, near Salamanca, New York, USA, May
American bobcat (Lynx rufus) male, drinking at water's edge, Texas, USA. April
Pair of Couch's spadefoot toads (Scaphiopus couchii) mating in water, Texas, USA. June
Painted bunting (Passerina ciris) male, perched on branch, Texas, USA. May
Vermilion flycatcher (Pyrocephalus obscurus) male, courtship display, Texas, USA. May
Male Baltimore oriole (Icterus galbula) foraging in flowering Crabapple (Malus sp)Male Baltimore oriole (Icterus galbula) foraging in flowering Crabapple (Malus sp.) tree in spring, Ithaca, New York, USA. May
Old male Espanola giant Galapagos tortoise (Chelonoidis hoodensis) stretching its neck in clearing, Las Tunas, Espanola Island, Galapagos Islands, Ecuador
Male Northern cardinal (Cardinalis cardinalis), in breeding plumage, perched in snow-covered Eastern redbud (Cercis canadensis) tree during spring snowstorm, Freeville, New York, USA. April
Mandarin duck (Aix galericulata) drake walking on snow-covered, frozen lake, London, UK. February
Black grouse (Tetrao tetrix) males fighting in snow during the breeding season, Viiksimo, Finland. April
Magnificent bird of paradise (Cicinnurus magnificus) male, leaning back and presenting his breast shield and raised cape to two females, courtship display, Arfak Mountains, West Papua, Indonesia
Costa's hummingbird (Calypte costae) male, portrait, showing purple iridescence of head and neck feathers, Coachella Valley, California, USA
Wilson's bird of paradise (Cicinnurus respublica) male, courtship display, Batanta Island, West Papua, Indonesia
Blue bird of paradise (Paradisaea rudolphi) male, perched on branch, Tari Valley vicinity, Southern Highlands Province, Papua New Guinea
Pair of Wilson's birds of paradise (Cicinnurus respublica), male performing courtship display to female from a small sapling in the middle of his display court, Batanta Island, West Papua