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Red deer (Cervus elaphus) stags running in herd across moorland, February
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April
Goldeneye (Bucephala clangula) male and two females on woodland loch at dawn on misty morning, Loch Vaa, Cairngorms National Park, Scotland, UK, April
Drake Mallard duck (Anas platyrhynchos) swimming through colourful reflections of trees at Glencoe Lochan. Scotland, UK. November
Iberian ibex (Capra pyrenaica), adult male, in snow covered landscape after snowfall. Sierra Nevada National Park, Andalusia, Spain. February
Whooper swans (Cygnus cygnus) pair engaged in courtship as snow falls, Myvatn lake, Iceland, March
Blackbird (Turdus merula) male eating winfall apples in snow, Broxwater, Cornwall, UK. December
Alpine ibex (Capra ibex) two adult males in mountain landscape. Alps, Aosta Valley, Gran Paradiso National Park, Italy. September
Great crested grebe (Podiceps cristatus) pair, courtship dance at dawn, Sierra de Grazalema Natural Park, Andalusia, Spain. April
Red deer (Cervus elaphus) stag bellowing, Richmond Park, London, England, UK. October
Saiga antelope (Saiga tatarica) male, standing in grassland in morning sunlight, Betpak Dala, Kazakhstan. Critically endangered
Caracal (Caracal caracal) male, standing on rock against blue sky, Spain. Captive, occurs in Africa and Asia
Western grey kangaroo (Macropus fuliginosus) pair, male standing and female lying down, Kangaroo Island, Australia
Three Common toads (Bufo bufo) in amplexus, Lucerne, Switzerland. March. Cropped
Common toad (Bufo bufo) and Common frog (Rana temporaria) in attempted amplexus, Lucerne, Switzerland. April
Yellow-bellied toad (Bombina variegata) male in pond, calling, Lucerne, Switzerland. July
Jumping spider (Hypositticus pubescens) male, resting on leaf, Lucerne, Switzerland. June. Focus stacked
Two Ibex (Capra ibex) males, standing on hillside, Lucerne, Switzerland. August
Rainforest toad (Incilius aucoinae) male, portrait, Osa Peninsula, Costa Rica
Two Iberian ibex (Capra pyrenaica), adult males, fighting. Sierra Nevada National Park, Andalusia, Spain. July
Iberian ibex (Capra pyrenaica), adult male, sitting down in snow covered landscape during snowfall. Sierra Nevada National Park, Andalusia, Spain. April
Bluethroat (Luscinia svecica cyanecula) male, perched on tree stump singing, Germany. March
Camargue horse (Equus ferus caballus) stallion rearing up. Camargue, France. June
Red deer (Cervus elaphus), young male, swimming through water. Bushy Park, London, UK. September
Wine throated hummingbird (Selasphorus ellioti) male, courtship display, Cloud forest, Atitlan Volcano, Solola, Guatemala
Blessed poison frog (Ranitomeya benedicta) male, carrying tadpoles on its back, transporting them to water, Yurimaguas, Alto Amazonas, Peru
Ruddy duck (Oxyura jamaicensis) male in courtship display. Valkenhorst nature reserve, Valkenswaard, The Netherlands. May
Walrus (Odobenus rosmarus) male resting in water, Kapp Lee, Svalbard, Norway, Arctic Ocean, July
Male Indigo bunting (Passerina cyanea) perched in flowering Serviceberry (Amelanchier) in spring. Salamanca, New York, USA. May
Two wild Onaqui stallions (Equus ferus) fighting for dominance in between their respective herd. Onaqui Mountain Herd Management Area, Utah, USA. September
Two wild Onaqui colts (Equus ferus) greeting one another. Onaqui Mountain Herd Management Area, Utah, USA. September
Humpback whales (Megaptera novaeangliae) pair engaging in courtship behavior, dancing for hours, off Tubuai, French Polynesia, Pacific Ocean
Two Cheetah (Acinonyx jubatus) brothers grooming one another. bonding behaviour, Linyanti, Chobe National Park, Botswana
Sika deer (Cervus nippon) stag wading through river in forest, Land of the Leopard National Park, Russian Far East. Taken with remote camera. August
Sika deer (Cervus nippon) stag walking on mountain slope at the edge of woodland, with forested mountains in the background, Land of the Leopard National Park, Russian Far East
Northern cardinal (Cardinalis cardinalis) male, perched on branch during snow storm, Milford, Connecticut, USA. January
American pit bull, male, standing on grass and autumn leaves, portrait, Haddam, Connecticut, USA. October
Adonis blue butterfly (Polyommatus bellargus) male, resting on Clover (Trifolium sp. ), Upper Bavaria, Germany. JuneAdonis blue butterfly (Polyommatus bellargus) male, resting on Clover (Trifolium sp.), Upper Bavaria, Germany. June
Lion (Panthera leo) male, juvenile, resting in tree during middle of the day to escape the heat, near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border, Tanzania
Vitelline masked weaver (Ploceus vitellinus) male, perched on hanging nest, Serengeti / Ngorongoro Conservation Area, Tanzania
Lion (Panthera leo) female, resting in tree during middle of day to escape the heat with juvenile male Lion approaching, near Ndutu, Ngorongoro Conservation Area / Serengeti National Park border
Lion (Panthera leo) male, scent marking its territory at dawn, Ngorongoro Conservation Area / Serengeti National Park boundary, northern Tanzania
Green-backed kingfisher (Actenoides monachus) male, perched on vine in forest understorey, Tangkoko National Park, northern Sulawesi, Indonesia
Puma (Puma concolor) male, walking through snow covered bushes, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Bluethroat (Luscinia svecica cyanecula) male, singing as a courtship display, Germany
Australian magpie (Gymnorhina tibicen) male perched on stump, bokeh effect, Canterbury, South Island, New Zealand
Black-naped monarch / Black-naped blue flycatcher (Hypothymis azurea) male, with insect prey in beak, feeding three chicks at nest, Wilpattu National Park, Sri Lanka
Magnificent frigatebirds (Fregata magnificens) pair in courtship display, male with gular sac inflated, North Seymour Island, Galapagos National Park, Galapagos Islands
