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Greater flamingos (Phoenicopterus ruber roseus) males displaying, lake Nakuru, Kenya
Male Arabian Leopard (Panthera pardus nimr) at the Arabian Wildlife Centre & captive-breeding project, Sharjah, United Arab Emirates
Flock of Stellers eiders (Polysticta stelleri) in flight, Batsfjord, Norway, March
Male King eider (Somateria spectabilis) displaying, Norway, February
Spruce grouse (Falcipennis canadensis) in forest, Denali National Park, Alaska, USA, September
RF - Polar Bear (Ursus maritimus) males fighting, Churchill, Canada, November
Polar bear (Ursus maritimus) curious young male on the newly frozen pack ice, Beaufort Sea, off the 1002 area of the Arctic National Wildlife Refuge, North Slope, Alaska
Stellers Eider (Polysticta stelleri) male swimming, Varangerfjord, Norway, April, Vulnerable species. Bookplate from Danny Greens The Long Journey North
Cavalier King Charles Spaniel, male puppy with tricolor coat, aged 3 months, at garden fence
Portrait of male Reindeer (Rangifer tarandus) with large antlers, in velvet. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, June
Red kangaroo male bounding towards camera. {Macropus rufus) NSW, Australia. The kangaroo is the animal long jump champion, able to clear 13m (the length of a bus) from a standing start
Booted racket tail (Ocreatus underwoodii) two males fighting in flight, Mindo, Pichincha, Ecuador
Lesser kestrel (Falco naumanni) male flying past during courtship with Mole cricket in its talons. Extremadura, Spain
Two male Lesser kestrels (Falco naumanni) in pursuit flight, one driving off the other from its nesting territory. Extremadura, Spain
Fleischmanns glassfrog (Hyalinobatrachium fleischmanni) male attending eggs on the underside of a leaf overhanging a rainforest stream
Red-eyed leaf frog (Agalychnis callidryas) male, Central Caribbean foothills, Costa Rica
Alpine Newt (Ichthyosaura / Triturus alpestris) male in mountain pool. Nordtirol, Austrian Alps. July
Four-spot orb weaver spider (Araneus quadratus) male approaching larger female to mate. Peak District National Park, Derbyshire, UK. October
Emerald damselflies (Lestes sponsa) mating, with male above. Peak District National Park, Derbyshire, UK. September
Sand lizard (Lacerta agilis) male basking. Dorset, England, UK, May
Brown hare (Lepus europaeus) adult male pursuing female, Derbyshire, UK, February
Red deer (Cervus elaphus) stag bellowing. Surrey, UK, October
Chaffinch (Fringilla coelebs) adult male drinking at garden pond, Scotland, UK. July
Kestrel (Falco tinnunculus) male, Hampstead Heath, England, UK
Lesser spotted woodpecker (Dryobates minor) male excavating nest hole in mature apple tree in ancient orchard. Herefordshire, UK. June
Chaffinch (Fringilla coelebs) male perched on mossy branch, Buckinghamshire, England, UK, February
Kingfisher (Alcedo atthis) male taking off from water after diving for fish, Worcestershire, England, UK, July
RF- Pair of Rock beauties (Holacanthus tricolor) spawning at dusk. The larger male rubs against the flank of female during the spawning rise. Cayman Islands, British West Indies. Caribbean Sea
Steller sealion (Eumetopias jubatus) male charging past the camera. Race Rocks, Victoria, Vancouver Island, British Columbia. Canada. North East Pacific Ocean
Portrait of a male Squarespot anthias (Pseudanthias pleurotaenia). Sangeang Island; Sumbawa, Indonesia. Flores Sea
Great white sharks (Carcharodon carcharias) males beneath the surface. Guadalupe Island, Baja California, Mexico. East Pacific Ocean
Blackbird (Turdus merula) male foraging on snow covered ground, during a blizzard. Derbyshire, UK. January (non-ex)
Ruppells weaver (Ploceus galbula) male building nest, Oman, May
Hourglass treefrog (Dendropsophus ebraccatus) male with inflated vocal sac calling at night, Central Caribbean foothills, Costa Rica
Three-striped poison frog (Ameerega trivittata) male carrying tadpoles, Panguana Reserve, Huanuco province, Amazon basin, Peru
Male Yellow warbler (Setophaga / Dendroica petechia) in breeding plumage singing amongst oak catkins and leaves, New York, USA, May
Male Bullethead parrotfish (Chlorurus sordidus) displaying in late afternoon. Ras Katy, Sinai, Egypt. Gulf of Aqaba, Red Sea
Male Scalefin anthias (Pseudanthias squamipinnis) swimming in open water, Ras Katy, Sharm El Sheikh, Sinai, Egypt. Gulf of Aqaba, Red Sea
Red deer (Cervus elaphus) stag bellowing during rut, Derbyshire, UK
Topi (Damaliscus korrigum / Damaliscus lunatus) territorial male standing on mound looking across savanna. Masai-Mara game reserve, Kenya
Greater sage-grouse (Centrocercus urophasianus) male displaying on a lek in snow, Sublette County, Wyoming, USA. April
Pine Grosbeak male (Pinicola enucleator)and traffic light Oulu, Finland, December
Mustang / wild horses, second year cremello colt Claro play fighting with bay, McCullough Peak herd, Wyoming, USA, July 2009
Rock beauty angelfish (Holacanthus tricolor) pair at dusk performing their courtship dance with the larger male circling the female, Georgetown, Grand Cayman, Cayman Islands, British West Indies
Adult male Hippopotamuses (Hippopotamus amphibius) posturing in agressive yawn behaviour, Luangwa River, South Luangwa National Park, Zambia, November. Digital composite (two frames stitched together)
Grizzly Bear (Ursus arctos horribilis) male in water. Coastal Katmai National Park, southwest Alaska, USA, August
Chinese Crested Dog, hairless bitch with puppy, male, 4 months
Male African Lion (Panthera leo) playing with 4 month cub. Big Marsh, near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania
