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Chinese Crested Dog, hairless bitch with puppy, male, 4 months
Male African Lion (Panthera leo) playing with 4 month cub. Big Marsh, near Ndutu, Nogorongoro Conservation Area, Serengeti National Park, Tanzania
Male Greenfinches (Carduelis chloris) squabbling in flight. Dorset, UK, March
Green crowned brilliant hummingbird (Heliodoxa jacula) male taking off to feed at feeder in garden, another waiting perched on lichen covered branch, Costa Rica, January
Emperor penguins (Aptenodytes forsteri) males dive through openings in the sea ice to take their first swim in over 4 months as they have spent the bleak winter incubating their eggs
Forester kangaroo (Macropus giganteus tasmaniensis) family group, male, female and large joey, Tasmania, Australia, February. Subspecies of Eastern grey kangaroo
A male Spiny Seahorse (Hippocampus guttulatus) hides in seagrass. Etang De Thau (Thau Lagoon), Montpellier, France, Europe, September
Chestnut arabian stallion running, California, USA
Grey Andalusian / Spanish stallion running, California, USA
Male Andean Cock-of-the-Rock (Rupicola peruvianus) displaying. Cock-of-the-Rock Lodge, Peru, August
A wild newborn colt (Equus caballus) standing up being nuzzled by its mother. Foothills of Reno, Nevada, USA, May
European Mouflon (Ovis musimon) male head profile. The Netherlands, January
Grey Wolf (Canis lupus) head portrait of male, in snow, Captive
Grey Wolf (Canis lupus) head portrait of male, sleeping in snow, Captive
British Shorthair tomcat, blue coated, portrait standing in straw with Pumpkins / Squash
Large adult male Leopard (Panthera pardus) walking through the bush at night, Sabi Sand Private Game Reserve, South Africa, June
Yellow-headed Blackbird (Xanthocephalus xanthocephalus) male calling and performing courtship / territorial display in cattail marsh, Mono Lake Basin, California, USA
Great Dane, black coated bitch, and Magyar Vizsla / Hungarian Pointer, tan coated maleGreat Dane, black coated bitch, and Magyar Vizsla /Hungarian Pointer, tan coated male, double portrait in profile together
Violet Sabrewing Hummingbird (Campylopterus hemileucurus) male hovering at feeder, Bosque de Paz, Costa Rica, Central America
African Elephant (Loxodonta africana) bulls in dominance fight, Mono Pools national Park, Zimbabwe, Southern Africa
A Shagya Arab stallion (Equus caballlus) is shown in hand in the courtyard of the Babolna Arabian Stud, Babolna, Komarom-Esztergom, Hungary
Portrait of male Boxer, light brindle coloured with natural ears, standing by lake shore, Illinois, USA
Weddell seal (Leptonychotes weddellii) male surfacing in ice hole, McMurdo Sound, Ross Sea, Antarctica, November 2008
Male Comet / Madagascar moon moth (Argema mittrei) recently emerged drying its wings. Andasibe-Mantadia National Park, Madagascar
Green woodpecker {Picus viridis} male in flight, Lorraine, France
Male Atlantic salmon {Salmo salar} leaping, migrating upstream to spawn, Perthshire, Scotland, UK, October
King Penguin (Aptenodytes patagonicus) adult standing amongst colony of chicks, Gold Harbor, South Georgia, South Atlantic Islands
RF - Head portrait of black Andalusian mare and stallion meeting for the first time in Spain, Europe
African lion (Panthera leo) male shaking water off mane after rain, Masai-Mara Game Reserve, Kenya. September
Polar bear (Ursus maritimus) young male playing / rolling around in the snow along a barrier island during autumn freeze up, Bernard Spit, 1002 area of the Arctic National Wildlife Refuge, Alaska
Common yellow throat (Geothlypis trichas) male, singing, Anchorage Provincial Park, Grand Manan Island, New Brunswick, Canada, June
Minute Glassfrog (Centrolene peristictum) male guarding eggs, Mindo, Pichincha, Ecuador, Vulnerable species
Male Bengal tiger (Panthera tigris tigris) known as Sundar (B2) patrolling territory, Bandhavgarh National Park, Madhya Pradesh, India. Endangered species
Tiger (Panthera tigris tigris), male portrait, Bandhavgarh, India, February 2013
RF- Rear view of male Bengal tiger (Panthera tigris tigris) walking on track. Bandhavgarh National Park, India. Endangered species
Humpback whale (Megaptera novaeangliae), fluke of female with male calf playing below, Vava'u, Tonga, Pacific Ocean
Kori bustard (Ardeotis kori) male, performing mating display, silhouetted at dusk, El Karama Wilderness Conservancy, northern Kenya
Iberian ibex (Capra pyrenaica), adult male, walking along mountain ridge. Sierra Nevada National Park, Andalusia, Spain. June
Goldeneye (Bucephala clangula) male on loch on misty morning, Loch Vaa, Cairngorms National Park, Scotland, UK, April
Silhouette of Red deer (Cervus elaphus) stag standing under a tree, Bradgate Park, Leicestershire, UK. October
Banded demoiselle dragonfly (Calopteryx splendens) male and female silhouetted, resting on reeds. Lower Tamar Lakes, Cornwall, England, UK. June
Kudu (Tragelaphus strepsiceros) male silhouetted, Kruger National Park, South Africa. Digital silhouette
Alpine ibex (Capra ibex) male in deep snow on a ridge during heavy snowfall, Gran Paradiso National Park, the Alps, Italy
RF - Puffins (Fratercula arctica) pair, resting on cliffside among flowering Sea thrift (Armeria maritima), Great Saltee Island, County Wexford, Republic of Ireland. May
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Japanese dwarf flying squirrel (Pteromys volans orii) male sitting in tree, portrait. Hokkaido, Japan. March
Red deer (Cervus elaphus) stags running in herd across moorland, February
Black Grouse (Tetrao tetrix) males at lek, Utajarvi, Finland, April