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RF- Grey Wolf (Canis lupus) howling at sunset (digitally enhanced)
Willow grouse / Willow Ptarmigan (Lagopus lagopus) in winter plumage. Wapusk National Park, Churchill, Manitoba, Canada, March
Giant Anteater (Myrmecophaga tridactyla) approaching livestock fence, Pantanal. Brazil
Owlfly sulphur (Libelloides coccajus) on flower, near Gavarnie, Pyrenees National Park, France, June
Leopard (Panthera pardus) resting in acacia tree, Samburu National Reserve, Kenya, Africa
Elephant trunk (Loxodonta africana), Etosha national park, Namibia, May
Hare, (Lepus europaeus) jumping in snowy field, Vosges, France, February
White stork (Ciconia ciconia) adult portrait, captive, Vogelpark Marlow, Germany, May
Lion (Panthera leo) male with bloody mane and flies, Masai-Mara Game Reserve, Kenya
Japanese Macaque (Macaca fuscata) young one, appearing quite relaxed in thermal pool, even falling asleep, in Jigokudani, Japan, February
Male Long-tailed duck (Clangula hyemalis), Batsfjord, Norway, March
Polar bear (Ursus maritimus) walking in snow, Svalbard, Norway, April
RF - Red fox (Vulpes vulpes) in dappled light on snow, Yellowstone National Park, USA. February
Red fox (Vulpes vulpes) walking in snow, Yellowstone National Park, USA, February
Red fox (Vulpes vulpes) hunting by pouncing onto prey through snow, Yellowstone National Park, USA, February
Lapwing (Vanellus vanellus), Skomer, Wales, UK. April 2011
Caribou / Reindeer bull (Rangifer tarandus) close-up grazing, Denali National Park, Alaska, USA, September
Atlantic Puffin (Fratercula arctica) two resting on a snow bank, Hornoya, Varanger, Norway, April
Leopard (Panthera pardus) female walking on fallen tree trunk, Samburu game reserve, Kenya
Hippo (Hippopotamus amphibius) male after a a mud bath, Masai-Mara Game Reserve, Kenya. Vulnerable species
Common seal (Phoca vitulina) pup hauled out on a beach at sunrise, Donna Nook Lincolnshire Wildlife Trust Reserve, Lincolnshire, England, UK, January
Grey seal (Halichoerus grypus) hauled out on a beach at sunrise, Donna Nook Lincolnshire Wildlife Trust Reserve, Lincolnshire, England, UK, January
Brown Bear (Ursus arctos) and reflection in pond. Finland, Europe, June
Large male Reindeer (Rangifer tarandus) standing in shallow waters of the Kronotsky lagoon, away from biting insects. Kronotsky Zapovednik Nature Reserve, Kamchatka Peninsula, Russian Far East, July
Australian sealion with young at sea surface {Neophoca cinerea} Australia. Endangered species
Apollo butterfly (Parnasius apollo) feeding on flower, Mount Terminillo, Rieti, Lazio, Italy, July 2011
Lion (Panthera leo) male using flehmen response to smell, Masai-Mara Game Reserve, Kenya. Vulnerable species
Chocoan green anole (Anolis parvauritus) on twig, Canande, Esmeraldas, Ecuador
Emerald glass frog (Espadarana prosoblepon) portrait, sitting on leaf, Buenaventura Reserve, Pinas, El Oro, Ecuador
Awa rocket frog (Hyloxalus awa) on stem, Rio Silanche, Pichincha, Ecuador, October 2013, Vulnerable species
Longnose harlequin frog (Atelopus sp) on leaf, Chinambi, Carchi, Ecuador
Lyre anole (Anolis lyra) on leaf, Chinambi, Carchi, Ecuador
Galapagos land iguana (Conolophus subcristatus) portrait, North Seymour Island, Galapagos, Vulnerable species
Giant river otter (Pteronura brasiliensis) yawning at water surface, Yasuni National Park, Orellana, Ecuador, Endangered species
African elephant (Loxodonta africana) walking, Brandberg, Namibia, Vulnerable species
Jocotoco antpittta (Grallaria ridgelyi) Tapichalaca Reserve, Loja, Ecuador
Sri Lankan Junglefowl (Gallus lafayettii) Yala National Park, Southern Province, Sri Lanka
Pinocchio rainfrog (Pristimantis appendiculatus) on a branch. Mindo, Pichincha, Ecuador
Horned dwarf iguana (Enyalioides praestabilis) on a branch. Sumaco National Park, Napo, Ecuador
White-throated kingfisher (Halcyon smyrnensis) Yala National Park, Southern Province, Sri Lanka
Carthusian Pink flower (Dianthus carthusianorum) detail of two stigmas, Nordtirol, Austrian Alps. June
Helmet Vanga (Euryceros prevostii) on nest incubating eggs. Masoala Peninsula National Park, north east Madagascar. IUCN Vulnerable Species
Grass snake (Natrix natrix) swimming amongst duck weed, Oxfordshire, UK
Greenfinch (Carduelis chloris) pair, one perched on branch and one hovering in snowfall, Derbyshire, February
Robin (Erithacus rubecula) on branch with Scarlet elfcup fungus (Sarcoscypha coccinea) spring. Dorset, UK, March
Bewicks swans (Cygnus columbianus) small flock in flight, Gloucestershire, UK February
Gannet (Morus bassanu) coming into land with stormy seas behind, Shetland Islands, Scotland, UK, September. Highly Commended in the Coastal and Marine category of the BWPA Awards 2015
Red grouse (Lagopus lagopus scotica) sitting in flowering heather, Peak District, UK, September
