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Salmon (Salmonidae) fry with egg case still attached, Lake Kuril, Kamchatka, Far East Russia, January
Red deer (Cervus elaphus) stag walking in snow covered pine forest, Cairngorms, Scotland, UK, December
Head portrait of grey Andalusian stallion in dark barn. Northern France, Europe
Boulengers tree frog (Rhacaphorus lateralis) recently rediscovered after 100 years. Coorg, Karnataka, India. Endemic to Western Ghats. Endangered species
Golden frog (Hylarana aurantiaca) with waterfall in background. Coorg, Karnataka, India
Malabar ramanella frog (Ramanella triangularis) Coorg Karnataka, India. Endemic to Western Ghats. Vulnerable species
Fairy pitta (Pitta nympha) in the forest in Guangshui, Hubei province, China. July
Red junglefowl (Gallus gallus) male bird walking through the forest of Hong Bung He, Dehong, Yunnan, China
Pallass squirrel (Callosciurus erythraeus) at Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China. April
Crested ibis (Nipponia nippon) flying in evening light, Yangxian Nature Reserve, Shaanxi, China. Endangered species. September
Wild yak, (Bos mutus), Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
Humpback whale (Megaptera novaeangliae) breaching - leaping out of the water, Sea of Cortez, Baja California, Mexico
Wood White butterfly (Leptidea sinapis) on Germander Speedwell (Veronica chamaedrys), Surrey, England, UK, May - Focus Stacked
Bullfinch (Pyrrhula pyrrhula) male walking along mossy twig, Buckinghamshire, England, UK, February
Cheetah (Acinonyx jubatus) walking near wildebeest, Masai Mara Nature Reserve, Kenya
Harebell carpenter bee (Chelostoma campanularum) at 4-5mm long one of Britains smallest bees, visiting Giant harebell (Campanula latifolia) to collect pollen, Pentwyn farm SSSI, Gwent Wildlife Trust
Cheetah (Acinonyx jubatus) male on the hunt with Crater rim in the background, Ngorongoro Crater, Tanzania
Reindeer (Rangifer tarandus) reindeer with bare antlers in rutting season, reintroduced Cairngorm Reindeer Herd, Cairngorm National Park, Speyside, Scotland, October
Lance-nosed chameleon (Calumma gallus), Andasibe-Mantadia National Park. Madagascar
Male banded demoiselle (Calopteryx splendens), resting on flower head, Lower Tamar Lakes, Cornwall, UK. June
RF - Common blue butterflies (Polyommatus icarus) resting on a backlit grass, evening light, Vealand Farm, Devon, UK. June
Polar bear (Ursus maritimus) walking in snow, Svalbard, Norway, April
Grey peacock-pheasant (Polyplectron bicalcaratum) walking through the forest at Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Raggy scorpionfish (Scorpaenopsis venosa) disputing territory. Ambon Bay, Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
RF - Bannerfish (Heniochus diphreutes) schooling in coral reef. Long exposure. North Male Atoll, Maldives. Indian Ocean
Robin (Erithacus rubecula) in the snow, Broxwater, Cornwall, UK. March
Four Mute swans (Cygnus olor) walking over snow, Hazerswoude, The Netherlands, February
Meadow brown butterfly (Maniola jurtina) on Common poppy seedhead (Papaver rhoeas), De Inslag, Brasschaat, Belgium August
Robin (Erithacus rubecula) on branch, Brasschaat, Belgium
Demoiselle cranes (Anthropoides virgo) flock, India
Cattle Egret (Bubulcus ibis) in breeding plumage, balancing on palm leaf, India
Woodlouse (Armadillidium pulchellum) Niedersechsische Elbtalaue Biosphere Reserve, Elbe Valley, Lower Saxony, Germany, December
Sanderling (Calidris alba) standing in shallow water, Nova Scotia, Canada, May
RF -Three Bison (Bison bison) walking through snow with herd feeding in background. Yellowstone National Park, Wyoming, USA. January
Grizzly cubs (Ursus arctos) two play fighting in the mist, near Brooks Falls in Katmai National Park, Alaska, USA, July
Red fox (Vulpes vulpes) resting and sitting on snow, Yellowstone National Park, USA, February
Coyote (Canis latrans) in wintry landscape, Yellowstone National Park, USA, February
Bison (Bison bison) standing in geothermal run-off in winter, Yellowstone National Park, Wyoming, USA, February
Red fox (Vulpes vulpes) in snow, Yellowstone National Park, Wyoming, USA. Wyoming, USA, January
Bison (Bison bison) standing steam from geothermal springs, Yellowstone National Park, Wyoming, USA, February
American oystercatcher (Haematopus palliatus) feeding, Fort Myers Beach, Gulf Coast, Florida, USA, March
Great egret (Ardea alba) in flight, Florida, USA, March
White Ibis (Eudocimus albus) feeding off Fort Myers beach, Gulf Coast, Florida, USA, March
RF- Red squirrel (Sciurus vulgaris) portrait, in snow, Cairngorms National Park, Scotland, March 2007
RF- Reindeer (Rangifer tarandus) portrait, semi-domesticated, small scale herding by local Sami, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden. September 2008
RF- Eurasian brown bear (Ursus arctos) mother walking with cub, Suomussalmi, Finland. July
John dory (Zeus faber) portrait, Babbacombe Bay, Devon, UK, October
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Portree, Skye, Inner Hebrides, Scotland, UK. June