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Thornbug (Umbonia crassicornis) San Jose, Costa Rica
Whooper swan (Cygnus cygnus) standing alone in snow storm, Hokkaido Japan February
European otter (Lutra lutra) with fish prey, with water splashing around, Kiskunsagi National Park, Hungary, July
Cocoi heron (Ardea cocoi) resting portrait, Pantanal, Brazil
Brimstone butterfly (Goneopteryx rhamni) male roosting on Cowslip (Primula veris) Bedfordshire, England, UK, April
Large copper butterfly (Lycaena dispar), female, Finland, July
Mazarine blue butterfly (Cyaniris semiargus) Aosta Valley, Gran Paradiso National Park, Italy
RF - Spicebush swallowtail butterfly (Papilio troilus) nectaring on Zinnia in farm garden, wild and free. Madison, Connecticut, USA
Meadow fritillary (Melitaea parthenoides) Riou de Meaulx, Provence, southern France, May
RF - Monarch butterfly (Danaus plexippus) with life-cycle tied to the presence of milkweed plants (Asclepias sp). East Haddam, Connecticut, USA. September
Marbled white butterfly (Melanargia galathea) resting on grass head, backlit against black background, Dunsdon, Devon, UK. July 2014
Common blue butterfly (Polyommatus icarus) resting on Scabious (Scabiosa colombaria) flower, Volehouse Farm, Devon, UK. August 2014
Duke of Burgundy butterfly (Hamearis lucina), Wiltshire, UK, June
Elephant Hawkmoth (Deilephila elpenor) Oxfordshire, UK. July
Chinese Sparrowhawk (Accipiter soloensis) flying Guangshui, Hubei province, China, July
Golden eagle (Aquila chrysaetos) male head portrait beak open calling, Kalvtrask, Vasterbotten, Sweden. December
Reindeer (Rangifer tarandus) in atmospheric light, herding in the Saarivuoma Sami community, at Jarama, Ovre Soppero, Lapland, Laponia, Norrbotten county, Sweden January
Greater kudu (Tragelaphus strepsiceros) female running, St Lucia Wetlands National Park, South Africa
Long-tailed duck (Clangula hyemalis) Batsfjord village harbour, Varanger Peninsula, Norway, March
Aquatic warbler (Acrocephalus paludicola) Nemunas River Delta, Lithuania. May. Vulnerable species species
Mute swan (Cygnus olor) in flight, Nemunas River Delta, Lithuania, May
Great white egret (Egretta alba or Ardea alba) catching cyprionid fish or carp in fish farm, Pusztaszer Protected Landscape, Kiskunsagi, Hungary, May
RF- Marine Iguana (Amblyrhynchus cristatus) head portrait, Fernandina island, Galapagos, Ecuador
RF- Ringed seal (Pusa hispida) hauled out on rock, Svalbard, Norway
Sword-billed hummingbird (Ensifera ensifera) hovering in flight, North-Ecuador
Ring-tailed cardinalfish (Ostorhinchus aureus), male protecting and incubating eggs in mouth, Philippines
Humpback whale (Megaptera novaeangliae), mother and calf swimming just below surface in front of the West Maui Mountains. South of Lahaina, West Maui, Hawaii
English springer spaniel standing in front of Rhododendron flowers. Haddam, Connecticut, USA. June
Giant Anteater (Myrmecophaga tridactyla), two sniffing air in savanna. Caiman Ecological Refuge, Southern Pantanal, Moto Grosso do Sul, Brazil
Snowy Egret (Egretta thula), Guerrero Negro, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Lyrtetail anthias (Pseudanthias squamipinnis) in water column with Peach anthias (Pseudanthias dispar) in background. Indonesia, October
Golden jackal (Canis aureus) walking on sand. Danube Delta, Romania, May
Little owl (Athene noctua) perched in wall. Danube Delta, Romania. May
Golden jackal (Canis aureus) howling in grassland. Danube Delta, Romania, May
Lion (Panthera leo), female amongst trees. Mana Pools National Park, Zimbabwe. November
African wild dog (Lycaon pictus), male and female play fighting. Mana Pools National Park, Zimbabwe
Willow tit (Poecile montanus) on snow, Kuusamo, Finland, March
Short-toed treecreeper (Certhia brachydactyla), Sierra de Grazalema Natural Park, Spain. July
Crested tit (Lophophanes cristatus) perched on lichen covered twig, Sierra de Grazalema Natural Park, southern Spain, July
Rufous-tailed scrub robin (Cercotrichas galactotes) perched on grape vine, Sevilla, Spain, August
Black egret (Egretta ardesiaca) forming wings into umbrella whilst fishing, Gambia
Polar bear (Ursus maritimus) walking along coast of Wrangel Island, Far Eastern Russia, October
Salmon (Salmonidae) fry with egg case still attached, Lake Kuril, Kamchatka, Far East Russia, January
Red deer (Cervus elaphus) stag walking in snow covered pine forest, Cairngorms, Scotland, UK, December
Head portrait of grey Andalusian stallion in dark barn. Northern France, Europe
Boulengers tree frog (Rhacaphorus lateralis) recently rediscovered after 100 years. Coorg, Karnataka, India. Endemic to Western Ghats. Endangered species
Golden frog (Hylarana aurantiaca) with waterfall in background. Coorg, Karnataka, India
