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Southern cassowary (Casuarius casuarius), female, walking through rainforest, Daintree lowlands, September
Comb-crested jacana (Irediparra gallinacea), Marrakai, Northern Territory, Australia, September
Hooded plover (Thinornis rubricollis) walking on beach looking for food. Skenes Creek, Victoria, Australia
Crested tit (Lophophanes cristatus), Sierra de Grazalema Natural Park, Cadiz, Andalusia, Southern Spain. December
Woodchat shrike (Lanius senator) perched on thistle, Sierra de Grazalema Natural Park, Cadiz, Andalusia, southern Spain, June
Ringneck blenny (Parablennius pilicornis) Vervece rock, Punta Campanella Marine Protected area, Costa Amalfitana / Amalfi coast, Italy, Tyrrhenian Sea, Mediterranean. October
Hippie anole (Anolis fraseri), Mindo, Ecuador
Flame-faced tanager (Tangara parzudakii), Ecuador
Ismenius tiger butterfly (Heliconius ismenius), Mindo, Ecuador
Portrait of a Hoatzin (Ophisthocomus hoazin) on black background. Yasuni National Park, Ecuador
Mandarin duck (Aix galericulata) male swimming with autumn colours reflected in the water, London, UK, November
Mallard (Anas platyrhynchos) male swimming with autumn colours reflected in the water, London, UK, November
Two Black-browed albatross (Thalassarche melanophris) rub their bills together - part of a bonding ritual. West Point Island, The Falkland Islands
Red-billed hornbill (Tockus erythrorhynchus) examines a pile elephant dung for a potential meal of small insects. Okavango Delta, Botswana
Kangaroo island echidna (Tachyglossus aculeatus multiaculeatus), a sub-species of the mainland species, crosses a dirt road. Kangaroo Island, South Australia, Australia. Endemic
Royal penguin (Eudyptes schlegeli) collects a pebble for its nest on the beach. Sandy bay, Macquarie Island, Australian Territory
Caribbean reef squid (Sepioteuthis sepioidea) portrait reflecting off the surface of the ocean at night, Eleuthera, Bahamas
RF - Maldive anemonefish (Amphiprion nigripes) swimming above their host Magnificent sea anemone (Heteractis magnifica) on coral reef
Magpie (Pica pica) perched on branch in snow, Lorraine, France, January
Red squirrel (Sciurus vulgaris) foraging on dry stone wall. Aigas Field Centre, Scottish Highlands. Scotland. October
Purple gallinule (Porphyrio martinicus) reaches for Giant bulrush seeds (Schoenoplectis californicus). Wakodahatchee Wetlands, Florida, USA, April
Canada goose (Branta canadensis) in flight in autumn, Maryland, USA. October
Wolf (Canis lupus), young male, portrait. Kiekinkoski, Finland. June
Eurasian brown bear (Ursus arctos) Kuikka, Kainuu, Finland, August
RF - Red squirrel (Sciurus vulgaris) feeding, sitting on branch. Cairngorms National Park, Scotland, UK. January. (This image may be licensed either as rights managed or royalty free.)
Saddle-billed stork (Ephippiorhynchus senegalensis) flapping wings after preening. Liuwa Plain National Park, Zambia
Taiwan Yellow tit, (Machlolophus holsti) Lu Shan, Taiwan endemic species
Swinhoes pheasant (Lophura swinhoii) male, endemic species, Yunlinshan, Taiwan
Dusky shark (Carcharhinus obscurus) portrait. Wild Coast, Eastern Cape, South Africa. June
Leopard shark (Stegostoma fasciatum) cruising along the edge of an underwater drop off, Philippines
Caracal (Caracal caracal) hunting, portrait. Namibia
Mute swan (Cygnus olor) backlit on a misty morning. Valkenhorst Nature reserve, Valkenswaard, The Netherlands, May
Tibetan antelope / Chiru (Pantholops hodgsonii) three silhouetted, Keke Xili / Hoh Xil Nature Reserve, Tibetan High plateau, Qinghai, China
Grey seal (Halichoerus grypus) female swims over oarweed (Laminaria digitata) in shallow water. Farne Islands, Northumberland, England, United Kingdom. North Sea
Little egret (Egretta garzetta) hunting in marshland, Suffolk. March
Atlantic puffin (Fratercula arctica) with beakful of fish, Saltee Islands, County Wexford, Republic of Ireland
Least Chipmunk (Tamias minimus) feeding on seeds, Grand Teton National Park, Wyoming, USA
Portrait of a Red-breasted nuthatch, (Sitta canadensis) with white background, Block island, Rhode Island, USA. Bird caught during scientific research
Red-necked tanager (Tangara cyanocephala) perched, Serra Bonita Private Natural Heritage Reserve (RPPN Serra Bonita), Camacan, Southern Bahia State, Eastern Brazil
Sperm whale (Physeter macrocephalus) mother and calf diving down, Faial Island, Azores, Portugal, Atlantic Ocean July
Crested guan (Penelope purpurascens), Finca Arroyo Negro, El Triunfo Biosphere Reserve, Chiapas, southern Mexico, April
Arabian partridge (Alectoris melanocephala) Al Mughsayl, Oman, November
Osprey (Pandion haliaetus) fishing. Acadia National Park, Maine, USA. May
North American river otter (Lontra canadensis) on snow covered bank, reflected in icy water. Yellowstone National Park, USA, January
Red fox (Vulpes vulpes) walking through deep winter snow. Hayden Valley, Yellowstone, USA. January
South Island tomtit (Petroica macrocephala macrocephala) male in snow. Arthurs Pass National Park, South Island, New Zealand. August
Striped Slender Robberflies (Leptogaster cylindrica) Ledston, Yorkshire, June. Focus stacked image
Marbled white butterfly (Melanargia galathea) roosting among tall grasses at sunrise and bathed in warm, early morning sunlight, Volehouse Moor, Devon, UK. July