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White Ibis (Eudocimus albus) feeding off Fort Myers beach, Gulf Coast, Florida, USA, March
RF- Red squirrel (Sciurus vulgaris) portrait, in snow, Cairngorms National Park, Scotland, March 2007
RF- Reindeer (Rangifer tarandus) portrait, semi-domesticated, small scale herding by local Sami, Sarek National Park, Laponia World Heritage Site, Lapland, Sweden. September 2008
RF- Eurasian brown bear (Ursus arctos) mother walking with cub, Suomussalmi, Finland. July
John dory (Zeus faber) portrait, Babbacombe Bay, Devon, UK, October
RF- White tailed sea eagle (Haliaeetus albicilla) in flight, Portree, Skye, Inner Hebrides, Scotland, UK. June
Humpback whale (Megaptera novaeangliae)just under surface, off Shetland, Scotland, UK. First ever underwater images of humpbacks in British waters
Dotterel (Charadrius morinellus) standing on rock, Cairngorms National Park, Scotland, UK, May
Whooper swan (Cygnus cygnus) pair on loch, Cairngorms National Park, Scotland, UK, November
Osprey (Pandion haliaetus) flying above water, Rothiemurchus, Cairngorms National Park, Scotland, UK, July
Red grouse (Lagopus lagopus scotica) in moorland, Cairngorms National Park, Grampian, Scotland, UK, April 2015
Ptarmigan (Lagopus mutus) male in snowy mountains, Cairngorms National Park, Scotland, UK, May 2015
Eurasian beaver (Castor fiber) foraging in loch at night, Knapdale, Argyll, Scotland, UK, June
Bottlenose dolphin (Tursiops truncatus) breaching, Moray Firth, Scotland, UK, July
Yellow stripe clingfish (Diademichthys lineatus) living in association with the Fire Urchin (Astropyga radiata) Lembeh Strait, North Sulawesi, Indonesia
Yellow-eyed penguin (Megadyptes antipodes) walking up a sand dune towards its nest. Otago Peninsula, Otago, South Island, New Zealand. February. Endangered Species
Juvenile Banggai cardinalfish (Pterapogon kauderni) with sea anemone (Heteractis magnifica). Lembeh Strait, North Sulawesi, Indonesia
American flamingo (Phoenicopterus ruber) at edge of water, Borrero Bay, Santa Cruz Island, Galapagos, Ecuador
Small heath butterfly (Coenonympha pamphilus) and snail, Ottange, Lorraine, France. August
Lesser spotted fritillary (Melitaea trivia) adult at rest on yellow flower, Bulgaria
Pine marten (Martes martes) leaping from branch, orange glow in sky behind from the lights of Inverness. Black Isle, Scotland, UK, November. Photographed by camera trap
Male impalas (Aepyceros melampus) standing in heavy rain, Masai-Mara Game Reserve, Kenya. February
Male black rhinoceros (Diceros bicornis) silhouetted against clouds, Masai-Mara Game Reserve, Kenya. March
Chestnut-headed Tesia (Tesia castaneocoronata) perched, Sichuan Province, China
Red-crowned Cranes (Grus japonensis) displaying and calling in snow, Hokkaido, Japan, February
Ptarmigan (Lagopus mutus) in winter plumage in flight, Utsjoki, Finland, April
Red-crowned Crane (Grus japonensis) in flight, Hokkaido, Japan, February
Great White Egret (Egretta alba) with water splashing from beak, Hungary January
Black Grouse (Tetrao tetrix) male displaying with breath condensing in cold air, Kuusamo, Finland April
Honey bee (Apis mellifera) in flight, Surrey, England, July
Female humpback whale (Megaptera novaeangliae) with its very young calf at the surface, Rurutu island, Australs Archipelago, French Polynesia. Pacific Ocean
Okavango robbers (Rhabdalestes maunensis) fish swimming up the temporary Selinda Spillway, northern Botswana. This shallow river cuts across the Kalahari Desert
Red lechwe (Kobus leche) silhouetted at sunrise, Okavango Delta, Botswana
Salvins albatross (Thalassarche salvini) flying at sea, closeup of head. Kaikoura, Canterbury, New Zealand, September. Vulnerable species
African elephant (Loxodonta africana) with Giraffes (Giraffa camelopardalis) and an Ostrich (Struthio camelus) in background. Etosha National Park, Namibia
Hooded crow (Corvus coronix) and Eurasian Magpie (Pica pica) waiting to scavenging White tailed eagles (Haliaeetus albicilla) food, Pusztazer, Hungary
Male Long tailed duck (Clangula hyemalis) on water, Batsfjord, Norway, March
Male King eider duck (Somateria spectabilis) on water, Batsfjord, Norway, March
Southern Tokoeka / Stewart Island Brown Kiwi (Apteryx australis lawryi) female drinking from stream after feeding on sand hoppers in rotting kelp. Ocean Beach, Stewart Island, New Zealand. Endemic
Small pearl bordered fritillary butterfly (Boloria selene) resting on grass, backlit underwing, Marsland Mouth, North Devon, UK, June
Marbled white butterfly (Melanargia galathea) resting on reed, Devon, UK, July
Graylag gosling (Anser anser) resting under parents wing, Pusztaszer, Hungary
Baby African elephant (Loxodonta africana), climbing up a riverbank, Chobe National Park, Botswana
Whooper Swan (Cygnus cygnus) male and female facing in opposite directions, central Finland, April
Three Marbled white butterflies (Melanargia galathea) resting on reeds, Devon, UK, July
Silver-studded blue butterfly (Plebejus argus) pair mating, resting on a Pyrimidal orchid (Anacamptis pyramidalis) flower, Gwithian Towans, Cornwall, UK, July
Burchells zebras (Equus quagga burchellii) standing side by side. Etosha NP, Namibia
Shining honeycreeper (Cyanerpes lucidus) Costa Rica