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Red Grouse (Lagopus lagopus scoticus) in Heather. Peak District, UK, August
Grizzly bear (Ursus arctos horribilis) walking, Yellowstone National Park, Wyoming, USA, October
Two Least Chipmunks (Tamias minimus) sitting on tree stump feeding, Yellowstone National Park, Wyoming, USA, October, crop of 1373882
Sperm Whale (Physeter macrocephalus) calf swimming uder its mother. Sao Miguel Island, Azores, July
Blue Shark (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Blue Sharks (Prionace glauca) at the sea surface. Santa Maria, Azores, August
Heath fritillary (Melitaea athalia) resting on scabious flower, Nordtirol, Austrian Alps, Austria, August
Comtois Horse (Equus caballus). Camargue reserve, Rhone, France, September
Male Whinchat (Saxicola rubetra) perched on Oil seed rape (Brassica napus) in a field, Vosges, France, April
Bridled nailtail wallaby (Onychogalea fraenata) Queensland, Australia, Endangered species, October
Marbled White Butterfly (Melanargia galathea) resting on grass stem, Devon, UK July. Winner of Seasons category of the BWPA 2011 competition
Red squirrel (Sciurus vulgaris) standing on its hind legs, Cairngorms National Park, Scotland, UK, March. Non-ex
Lapwing (Vanellus vanellus) group roosting in the lagoons at Titchwell RSPB Reserve, Norfolk, UK, October. Non-ex
Black-Winged Stilt (Himantopus himantopus) wading. Etosha National Park, Namibia, January
Jaguar (Panthera onca palustris) male wading through water at the shore of the Piquiri River. The Pantanal wetlands of Mato Grosso, Mato Grosso State, Brazil, October
Edible chanterelle fungus (Cantharellus cibarius) showing spore dispersal pattern over 24 hours on black card
Pheasant (Phasianus colchicus) male amongst Bluebell flowers, UK, April
A male Leafy Seadragon (Phycodurus eques) swimming. Wool Bay Jetty, Edithburgh, Yorke Peninsular, South Australia, November
A juvenile Weedy Seadragon (Phyllopteryx taeniolatus) in its habitat. Flinders Jetty, Melbourne, Victoria, Australia, March
A young Spiny / Yellow / Longsnouted Seahorse (Hippocampus guttulatus), in an aquarium attached to plastic seagrass leaves. This seahorse is part of a UK seahorse breeding program
West Highland White Terrier leaping
Long Water Measurer (Hydrometra stagnorum) drinking by piercing surface tension with mouthparts. Captive, UK, July
Couchs Spadefoot (Scaphiopus couchii) adult at night calling with vocal sac inflated. Laredo, Webb County, South Texas, USA, April
Red Fox (Vulpes vulpes) pouncing in grass. Vosges, France, July
Red Fox (Vulpes vulpes) resting on straw bale in field. Vosges, France, July
Roe Deer (Capreolus capreolus) swimming across River Allier, France, February
Redpoll (Carduelis flammea) perched on a branch in snow. Breton Marsh, French Atlantic Coast, March
Yellow Labrador Retriever standing in glasswort against a blue sky. Monterey Peninsula, California, USA, February
Portrait of a wild Horse (Equus caballus). The foothills of southwest Reno, Nevada, USA, May
Two Brown bears (Ursus arctos) fighting in woodland wetlands, Kuhmo, Finland, July
Mute Swan (Cygnus olor) standing on frosty grass. The Netherlands, March
Atlantic salmon (Salmo salar) moving upriver to spawn. Lligwy River nr Betws y Coed, Gwynedd, North Wales, UK, November
Brown bear (Ursus arctos) standing beside lake looking for salmon, Kamchatka, Far east Russia, August
African elephant (Loxodonta africana) at waterhole with Giraffe in the background, Etosha National Park, Namibia, August
European hare (Lepus europaeus) mating pair boxing in field, Slovakia
Panther Chameleon {Furcifer pardalis} walking along branch on coast, Masoala Peninsula National Park, north east Madagascar
Common / Eurasian cranes (Grus grus) small flock of juveniles taking flight over the frozen, snow covered pastureland on a foggy winter morning
Dipper (Cinclus cinclus) landing on exposed stone, with insect prey for chicks in beak, Brecon Beacons National Park, Wales, UK
Sri Lankan Leopard (Panthera pardus kotiya) soft focus running, captive
Small flock of Greater flamingo (Phoenicopterus roseus) wading on inland Kalloni lake, Lesbos, Greece
Silhouette of Black-winged stilt (Himantopus himantopus) standing on exposed rock over water, with reflections, Lesbos, Greece
African lion (Panthera leo) male and female at sunrise, Masai Mara National Reserve, Kenya, January
Red grouse {Lagopus lagopus scoticus} on heather moorland, Derwent Edge, Peak District National Park, UK, September
European Lynx (Lynx lynx) in birch forest in snow, Tromso, Norway, captive, April
Red Squirrel (Sciurus vulgaris) leaping through Ash Tree in deciduous woodland, Berwickshire, Scotland, UK, February
Grizzly bear (Ursus arctos horribilis) catching salmon in Brooks river, Katmai National Park, Alaska, USA, July
RF - Chinstrap Penguins (Pygoscelis antarcticus) once calling, South Shetland Islands, Antarctica Continent
Erect-crested penguins (Eudyptes sclateri) pair on coast, Proclamation Island, Bounty Islands, New Zealand Sub-Antarctic Islands. Endemic to Antipodes and Bounty Islands. Endangered species
