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Flightless cormorant (Phalacrocorax harrisi) coming ashore amongst crabs. Clearwater Bay, Isabela Island, Galapagos. Sequence 3 of 3
Red fox (Vulpes vulpes) cub at the entrance to the den on allotment, North London, England. April 2020
Anolis lizard (Anolis sp. ) male displaying in Corcovado National Park, Costa RicaAnolis lizard (Anolis sp.) male displaying in Corcovado National Park, Costa Rica
Caribbean reef squid (Sepioteuthis sepioidea) portrait reflecting off the surface of the ocean at night, Eleuthera, Bahamas
Galapagos penguin (Spheniscus mendiculus) hunting off Bartolome Islet, Santiago Island, Galapagos, Ecuador. December
Copper-cheeked frog (Chalcorana labialis) from riparian lowland rainforest on the Cross-island trail, Tioman Island, Malaysia
Potato grouper (Epinephelus tukala) in Passe Dubois / Dubois channel, Aldabra, Indian Ocean
RF-Polar bear (Ursus maritimus) swimming in Beaufort Sea, portrait. Near Kaktovik, Arctic National Wildlife Refuge, Alaska, USA. September
Corncrake (Crex crex) calling at dawn on machair, North Uist, Outer Hebrides, Scotland, UK, June
Dipper (Cinclus cinclus) in river, Finland, March
Red-throated Diver (Gavia stellata) portrait, Iceland, June
Hazel Grouse (Tetrastes bonasia), Kuusamo, Finland, December
Northern wheatear (Oenanthe oenanthe) Arcos de la Frontera, southern Spain. April
Mediterranean gull (Larus melanocephalus) in summer plumage, Hayling Island, Hampshire, England, UK, July
Great horned owl (Bubo virginianus) in flight. Saskatchewan, Canada, August
Grey shrike (Lanius meridionalis) perched, Navarra province. Spain, July
Coyote (Canis latrans) foraging in snow. Hayden Valley, Yellowstone National Park, USA. January 2019
RF - Male African lion (Panthera leo) scratching tree at Big Marsh, Nogorongoro Conservation Area / Serengeti National Park, Tanzania. March
Male Anole Lizards (Anolis sp) in territorial display to each other. Rainforest canopy, Rio Claro Reserve, Magdalena Valley, Colombia, South America
Atlantic walruses (Odobenus rosmarus) resting on ice, with two large individuals facing off just before the one on the right stabbed the other with his tusks, Svalbard, Norway, June
Short-snouted seahorse (Hippocampus hippocampus) portrait. Tenerife, Canary Islands
Mandarin duck (Aix galericulata) reflected in water. Richmond Park, London, England, UK
American quarter horse standing in forest, portrait. Alberta, Canada. February
Chinese goral (Naemorhedus griseus) in snow. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Cheetah (Acinonyx jubatus) walking, silhouetted at dusk. Zimanga private game reserve, KwaZulu-Natal, South Africa
African elephants (Loxodonta africana) herd silhouetted at sunset. Chobe National Park, Botswana
Ruwenzori side-striped chameleon, (Trioceros rudis), hanging from fern, Nyungwe Forest NP, Rwanda, December. Non-ex
Common wasp (Vespula vulgaris) feeding on bread and jam. Monmouthshire, Wales, UK. March
Silky shark (Carcharhinus falciformis), three in Caribbean Sea off Gardens of the Queen National Park, Cuba
Red deer (Cervus elaphus) stags in snowy Pine forest. Scotland, UK. March
Giant panda (Ailuropoda melanoleuca) portrait, in falling snow. Captive bred, in enclosure, Panda Breeding Centre, Wolong, Sichuan Province, China. February 2008
Coffee bee hawk moth (Cephaonodes hylas) Amagasaki City, Hyogo, Japan
African Spoonbill (Platalea alba) preening, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Many-coloured rush tyrant (Tachuris rubrigastra) perched on reed stem at the waters edge. Lake Titicaca, Puno, Peru. September. Cropped
Great egret (Ardea alba) with prey in beak, in evening light. Myakka River State Park, Florida, USA, March. Sequence 2/2
Tricoloured heron (Egretta tricolor) fishing by flying low over water, amongst Water lilies. Everglades National Park, Florida, USA. March
Onager (Equus hemionus), Negev desert, Israel, April
Sand-loving / Square-headed wasp (Tachysphex costae) feeding on Sea holly flowers (Eryngium maritimum) on a beach, Sardinia, Italy, June
Blue-headed quail-dove (Starnoenas cyanocephala) profile, Cuba. Endangered species. Endemic
Espanola mockingbird (Mimus macdonaldi) on beach, Galapagos
Purple gallinule (Porphyrio martinicus) reaches for giant bulrush (Schoenoplectis californicus) seeds. Wakodahatchee Wetlands, Florida, USA, April
Thorny or spiny seahorse (Hippocampus histrix) portrait made off Anilao, Philippines
RF - Roe buck (Capreolus capreolus) stood in rough grassland in evening light, Scotland. September
Pectoral sandpiper (Calidris melanotos) juvenile, migrant, Llanrhystud, Ceredigion, West Wales
Pine marten (Martes martes) walking along edge of water, orange glow in sky behind from the lights of Inverness. Black Isle, Scotland, UK, September. Photographed by camera trap
Beetle, (Rhagonycha fulva), feeding on oxeye daisy flower, Scotland, UK, July
Red Fox (Vulpes vulpes) watching prey through long grass. London, UK. November
Intermediate egret (Egrettta intermedia), Keoladeo NP, Bharatpur, India