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Whale shark (Rhincodon typus) pectoral fin and skin markings detail, Triton Bay, West Papua, Indonesia, Pacific Ocean
Bargibants pygmy seahorse (Hippocampus bargibanti) camouflaged in a seafan (Muricella sp)Bargibants pygmy seahorse (Hippocampus bargibanti) camouflaged in a seafan (Muricella sp.), Raja Ampat, West Papua, Indonesia, Pacific Ocean
Male Dendtric jawfish (Opistognathus dendriticus) with incubating eggs in his mouth, Dampier Strait, Raja Ampat, Indonesia, Pacific Ocean
Tassled scorpionfish (Scorpaenopsis oxycephala), portrait, Triton Bay, West Papua, Indonesia, Pacific Ocean
Emu (Dromaius novaehollandiae) drinking at freshwater pool, Biloela, Queensland, Australia
Pair of Metallic starlings (Aplonis metallica) collecting nesting material, Tully Falls, Queensland, Australia
Female Eclectus parrot (Eclectus roratus) feeding in rainforest, Cape York, Queensland, Australia
Female Plum-headed finch (Neochima modesta) feeding in long grass, Dalby, Queensland, Australia
Mareeba rock wallaby (Petrogale mareeba) standing on granite boulders, Atherton Tablelands, Queensland, Australia
Male Numbat (Myrmecobius fasciatus) in dry woodland, Dryandra woodland, Western Australia
White-lipped green tree frog (Litoria infrafrenata) clinging on to a branch, Etty Bay, Wet Tropics World Heritage area, Queensland, Australia
Common mist frog (Litoria rheocola) sitting on a leaf at night, Wet Tropics World Heritage area, Queensland, Australia
Mouse-tailed bat (Rhinopoma hardwickii), portrait, Sinai Penimsula, Egypt
Northern freetail bat (Chaerophon jobensis), portrait, Mt Isa, Queensland, Australia
Large-eared horseshoe bat (Rhinolophus robertsi), portrait, Atherton, Queensland, Australia
Dippers (Cinclus cinclus) standing on ice, ready to fight over feeding territory, Kuusamo, Finland, February
Firecrest (Regulus ignicapilla) amongst oak leaves in autumn, Parainen Uto, Finland. October
Chocolate border collie (Canis familiaris) playing in water, Maryland, USA. October
Brown bear (Ursus arctos), portrait, Kronotsky Nature Reserve, Kamchatka, Russia
Male Saiga antelope (Saiga tatarica) rutting in winter, The Black Lands National Park, Kalmykia, Russia
Brown noddy (Anous stolidus) perching on rock face, Isabela Island, Galapagos, South America
Pair of Waved albatross (Phoebastria irrorata) courtship, Espanola Island, Galapagos, South America
Waved albatross (Phoebastria irrorata), pair, Espanola Island, Galapagos, South America
Sierra Negra giant tortoise (Chelonoidis guntheri) walking, Isabela Island, Galapagos, South America
Male Santiago lava-lizard (Microlophus jacobi), among Galapagos carpetweed (Sesuvium edmonstoni), Bainbridge Islets, Galapagos, South America
Santa Fe land iguana (Conolophus pallidus) with Giant prickly-pear cactus (Opuntia echios), Santa Fe Island, Galapagos, South America
San Cristobal lava lizard (Microlophus bivittatus), San Cristobal Island, Galapagos, South America
Lava heron (Butorides sundevalli), fishing from rocks, Santiago Island, Galapagos, South America
Galapagos sea lion (Zalophus wollebaeki) mother with newborn pup sitting on beach, Espanola Island, Galapagos, South America
Galapagos land iguana (Conolophus subcristatus) recently reintroduced to Santiago island where it had become extinct, feeding on Yellow cordia (Cordia lutea) leaves, Santiago Island, Galapagos
Darwins woodpecker finch (Cactospiza pallida) using twig as tool to extract beetle grub from dead branch, Santa Cruz Island, Galapagos, South America
Darwins small ground finch (Geospiza fuliginosa), inspecting Galapagos giant tortoise for ticks, Isabela Island, Galapagos, South America
Darwins medium ground finch (Geospiza fortis) feeding on wild seeds (Tournefortia sp), Santa Cruz Island, Galapagos, South America
Close-up of Eurasian lynx (Lynx lynx) kitten, aged eight months, face, breeding and reintroduction program, Germany. Captive. January
Two juvenile Asian elephants (Elephas maximus) having fun bathing and playing in river, Pairi Daiza, Belgium. Captive
Dunnock (Prunella modularis) perching on tree branch, Belgium. July
Close up of a Caribbean flamingo (Phoenicopterus ruber) in the breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Bookplate
Caribbean flamingo (Phoenicopterus ruber) flock, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June. Bookplate
Mute swan (Cygnus olor) landing on frozen lake, Richmond Park, London, UK. February
Female Fox (Vulpes vulpes) caught in a shaft of light on the porch of a farmhouse, Verteskozma, Hungary
Female Fox (Vulpes vulpes) walking on tree trunk in heavy snowstorm in the dark, Verteskozma, Hungary
Common tern (Sterna hirundo) catching a mayfly, Hjalstaviken, Uppland, Sweden. Nordic Nature Photo Competition 2021 Highly Commended - Nordic Bird Photo of the Year category
Male Bleekers parrotfish (Chlorurus bleekeri) races across a coral reef as it displays to females.Yanggefo Island, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Crescent-tail bigeye (Priacanthus hamrur) school on a coral reef, shot with a long exposure. North Male Atoll, Maldives. Indian Ocean
Male Fifteen spined stickleback / Sea stickleback (Spinachia spinachia) guarding a nest in Spaghetti seaweed / Thong weed (Himanthalia elongata), Isle of Coll, Inner Hebrides, Scotland, UK
Common dolphin (Delphinus delphis) porpoising at the surface to breathe, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Common dolphin (Delphinus delphis) leaping out of sea, Isle of Coll, Inner Hebrides, Scotland, UK, Atlantic Ocean. August
Male Lumpsucker (Cyclopterus lumpus) swimming through Kelp forest (Laminaria hyperborea), Loch Carron, Highlands, Scotland, UK