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Brazilian tapir (Tapirus terrestris) portrait, Tambopata Research Centre, Peru
Guianan cock-of-the-rock (Rupicola rupicola), two males fighting during lek, Guianan Shield, Suriname
Striped Dolphin (Stenella coeruleoalba) in the strait of Gibraltar, Spain
Southern elephant seal (Mirounga leonina), fight between two males, Saint Andrew, South Georgia
Komodo dragon (Varanus komodoensis) Rinca Island, Indonesia
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
Nurse sharks (Ginglymostoma cirratum) over seagrass at night. White spiracle behind eye is visible. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Sol de Manana geyser bassin, Reserva Eduardo Avaroa, Altiplano, Bolivia
Puna / James flamingo (Phoenicoparrus jamesi) flock on Laguna Colorado, Reserva Eduardo Avaroa, Altiplano, Bolivia
Egyptian goose (Alopochen aegyptiaca) pair amongst Devil s-thorn yellow flowers (Tribulus terrestris) Etosha National Park, Namibia, March
Scimitar-horned oryx (Oryx dammah) captive in enclosure of Souss Massa National Park, Morocco. Extinct in the Wild
Chamois (Rupicapra rupicapra) in the snow, Jura Switzerland, November
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Seychelles kestrel (Falco araeus), adult perched on wire, Republic of Seychelles Vulnerable species
Hippopotamus (Hippopotamus Amphibius) territorial aggressive behavior, Chobe National Park, Botswana
Sea apple (Pseudocolochirus violaceus) Komodo National Park, Indonesia
Thorny dragon (Moloch horridus) in desert habitat, Australia
Banded darter (Sympetrum pedemontanum) resting on flower, Netherlands
Typical landscape in Tuscany, with the farmhouse Belvedere near to San Quirico d Orcia, Italy, June
Toque Macaque (Macaca sinica) baby peering through mothers legs, Yala National Park, Southern Province, Sri Lanka
Rock formations at Brimham Rocks created by variable erosion of soft and hard layers of carboniferous age millstone grit, Harrogate, Yorkshire, England, UK, October 2016
School of Munks devil ray (Mobula munkiana), feeding on plankton at night, photographed on a long exposure, Espiritu Santo Island, Sea of Cortez, Baja California, Mexico, East Pacific Ocean
Strangler fig (Ficus sp) growing over Hoop Pine tree(Araucaria cunninghamii) with branches knotting round each other, Morans Falls Track, Green Mountains
Sumatran orangutan (Pongo abelii) female next to infant, Gunung Leuser National Park, Indonesia, UNESCO World Heritage site, November
European ground squirrels / Sousliks (Spermophilus citellus) in grass with daisies, Gerasdorf, Austria. April
Sumatran orangutan (Pongo abelii) female moving between lianas, Gunung Leuser National Park, UNESCO World Heritage site, November
African elephant (Loxodonta africana) feeding on grasses, Chobe River, Chobe National Park, Botswana. May
Black korhaan (Eupodotis afra) male displaying and calling. Etosha National Park, Namibia, March
African buffalo (Syncerus caffer) feeding on water lillies in Chobe River, Chobe National Park, Botswana. May
Indian rhinoceros (Rhinoceros unicornis) portrait. Manas National Park UNESCO World Heritage Site, Assam, India
Honey bee (Apis mellifera) on comb with honey, Kiel, Germany, June
Banyan tree (Ficus macrophylla) Valley of the Shadows, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Dingo (Canis lupus dingo) on the beach at night, Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Birch tree (Betula) growing in Llyn Padarn, Llanberis, Snowdonia National Park, North Wales, UK, March
Indian hog deer (Hyelaphus porcinus) mother and fawn, Manas National Park UNESCO World Heritage Site, Assam, India
Blackbuck (Antelope cervicapra), male, Tal Chhapar Wildlife Sanctuary, Rajasthan, India
Jaguar (Panthera onca) male and female four month old cubs playing, native to Southern and Central America, captive
Jaguar (Panthera onca) mother grooming four month cub, native to Southern and Central America, captive
Jaguar (Panthera onca) female snarling, native to Southern and Central America, captive
Madagascar crested wood ibis (Lophotibis cristata) preening, endemic to dry forest of Madagascar, captive
Llama standing in mud at the edge of Laguna Colorada, Bolivia
Giant cushion plant (Azorella compacta). Bolivia
Brindle-and-white Lurcher puppy, age 8 weeks, with crossed paws
High Altiplano with tussock grass called Paja brava (Festuca orthophylla) showing clonal growth spread. Bolivia. December 2016
English Oak (Quercus robur) in arable field with wheat, composite image showing all four seasons, Surrey, England, UK
Rolling farmland in the Overberg region near Villiersdorp, Western Cape, South Africa. December 2014
Winter storm on the coast near Arnastapi (rock arch of Gatklettur), Snaefellsness Peninsula, Iceland, February 2016
Border collie dog sitting with the church at Bjarnarhofn in background, Snaefellsness Peninsula, Iceland, March 2015