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Green hydra (Hydra viridissima) found in a garden pond. Derbyshire, UK. September. Winner of the Hidden Britain Category of the British Wildlife Photographer of the Year Awards (BWPA) Competition 2017
Harvest mouse (Micromys minutus) climbing on frosty seedhead, Hertfordshire, England, UK, January, Controlled conditions
Harvest mice (Micromys minutus) on grass stems, Devon, UK. July 2016. Captive
Silver-studded blue butterflies (Plebejus argus) resting on grass and backlit by the sun, Upton Towans, Cornwall, UK. June 2017
Common blue butterflies (Polyommatus icarus) resting on grasses, Vealand Farm, Devon, UK. June 2017
Mountain hare (Lepus timidus) grooming itself, with back foot raised, Cairngorms National Park, Scotland, UK, February. Highly Commended in the Animal Behaviour category of the British Wildlife
Striped mackerel (Rastrelliger kanagurta) mouth wide open as it swims through the water, filtering zoopankton with its gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Great hammerhead shark (Sphyrna mokarran) in shallow water. South Bimini, Bahamas. The Bahamas National Shark Sanctuary. Gulf Stream, West Atlantic Ocean
Banded demoiselle (Calopteryx splendens) male close up detail of head and thorax, Hertfordshire, England, UK, June. Captive. Focus stacked
Eagle fern (Pteridium aquilinum) frond unfurling, Montseny Natural Park, a UNESCO National Park and Biosphere Reserve, Barcelona, Catalonia, Spain, June 2016
Emerald Glass Frog (Centrolenella proseblepan) on leaf. Mid-altitude rainforest, Bosque de Paz, Pacific slope, Costa Rica, Central America
Green turtle (Chelonia mydas) female on a coral reef. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Strawberry poison dart frog (Oophaga pumilio) La Selva Field Station, Costa Rica
Strawberry poison dart frog (Oophaga / Dendrobates pumilio) sitting in cup fungus, La Selva Field Station, Costa Rica
Forest with mist on the slopes of Mount Lidgbird. Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia
Parsons Chameleon (Calumma parsonii) climbing in rainforest understorey. Andasibe-Mantadia National Park, eastern Madagascar
Deadvlei with dead Camel thorn trees, Namib-Naukluft National Park, Namib Desert, Namibia
Northern rockhopper penguin (Eudyptes moseleyi) portrait, Gough Island, Gough and Inaccessible Islands UNESCO World Heritage Site, South Atlantic
Track running through woodland with Bluebells (Hyacinthoides non-scripta) flowering, near Cerne Abbas, Dorset, England, UK, April 2014
Sau and Vilanova de Sau, viewed from Tavertet and Santa Maria del Corco, Les Guilleries Natural Park, Barcelona, Catalonia, June 2016
Lago di Landro at dawn, Dolomite Mountains, Sud Tirol / Alto Adige, ItalyLago di Landro at dawn, Dolomite Mountains, Sud Tirol/Alto Adige, Italy
Green-winged macaws (Ara chloroptera) preening each other. Brazil. South America
Peacock butterflies (Inachis io) feeding on corn marigolds (Glebionis segetum) on agricultural headland. England, UK
Fringed Tulip Dallas (Pink) and Tulipa Swan Wings (White) and Tulipa Valery Gergiev (red) growing in garden border. England, UK
Spring crocus (Crocus vernus) flowering en masse on the Campo Imperatore, Abruzzo, Italy. April
Black bear cubs (Ursus americanus) standing in a tree, Minnesota, USA, June
Pod of Risso Dolphin (Grampus griseus) at surface, Baja California, Mexico
Spinner dolphin (Stenella longirostris) engaged in spinning manoeuvre, Sri Lanka. Composite sequence of images
Reflections of clouds and landscape in Loch Arkaig, Glen Dessary, Lochaber, Scotland, UK, September 2016
Waterlilies (Nymphaea alba) underwater in lake. Alps, Ain, France
Fireflies (Luciola parvula himebotaru) flashing at night for courtship and reproduction. Gifu, Japan. Composite image
Mountain view towards Loch Nevis overlooking Knoydart peninsula with Skye beyond. Taken from Sgurr Cos na Breachd-laoidh, Lochaber, Scotland, UK, September 2016
Caribbean manatee or West Indian manatee baby, age two days, (Trichechus manatus) captive, Beauval Zoo, France
Caribbean manatee or West Indian manatee (Trichechus manatus) mother with baby, age two days, captive, Beauval Zoo, France
Roe deer (Capreolus capreolus) young male walking through snow. Close-up, portrait. Norway, February
Adult Kea (Nestor notabilis) close-up portrait. Arthurs Pass National Park, New Zealand, July. Endangered Species
Crocodile monitor (Varanus salvadorii) portrait, captive, occurs in New Guinea
Wood sandpiper (Tringa glareola) in reed bed, Finland, May
Long-tailed tit (Aegithalos caudatus caudatus) placing feather in nest, Finland. April
Blackcap (Sylvia atricapilla) female and two males huddling together for warmth before going to sleep, Finland, May
Ural owl (Strix uralensis), juvenile, Finland, June
Bearded reedling / tit (Panurus biarmicus), male in snow, Finland, March
Lesser spotted woodpecker (Dendrocopos minor), male at nest hole, Finland, June
Pearl-bordered fritillary (Boloria euphrosyne) feeding from Wild Rosemary (Rhododendron tomentosum), Finland, June
Lesser purple emperor butterfly (Apatura ilia), sitting on a leaf, Finland, August
Orange-tip (Anthocharis cardamines), male feeding from Wood Cranesbill (Geranium sylvaticum), Finland, June
Caribbean Flamingo (Phoenicopterus ruber) chick age 5 days, walking around nest, breeding colony, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, June
Suspension bridge across the river, entrance to Gunung Mulu National Park UNESCO Natural World Heritage Site, Malaysian Borneo