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Millstones made from millstone grit beneath Stannage Edge, Hathersage, Derbyshire, November 2016
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June
Galapagos sea lion (Zalophus wollebaeki) pup on shore, Galapagos
Deserta Grande wolf spider (Hogna ingens), Deserta Grande, Madeira, Portugal. Critically endangered
Giant panda (Ailuropoda melanoleuca) female, Huan Huan, holding baby age three months, Beauval Zoo, France, November 2017
Deep sea jellyfish (Periphylla periphylla), Trondheimsfjord, Norway, July
Fin whale (Balaenoptera physalus) just below surface, south Barcelona coast, Spain, Mediterranean Sea
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Fin whale (Balaenoptera physalus) with ctenophore in front of its mouth, south Barcelona coast, Spain, Mediterranean Sea
Short-finned pilot whale (Globicephala macrorhynchus) showing underside whilst vertical underwater, South Tenerife, Canary Islands, Atlantic, October
Silver tabby cat, Loki, age 7 months, walking with tail up
Norfolk Terrier dog, Otto, age 1 year
Border Terrier puppy, age 5 weeks, with raised paw
Giant sequoia (Sequoiadendron giganteum) trees in Sequoia National Park, California, USA, September 2014
Conifer trees in mist at Alpe de Lerosa, Dolomite Mountains, Belluno Province, Veneto, Italy, September 2015
Thomass Langur (Presbytis thomasi), animal endemic to northern Sumatra. These primates have relatively long arms and legs, adapted for an acrobatic life in the crowns of trees
Bairds tapir (Tapirus bairdii) Corcovado National Park, Costa Rica, May. Endangered
Stellers Sea Eagle (Haliaeetus pelagicus) with prey in beak in mid-air, Hokkaido, Japan, February
Close up if a Great hammerhead shark (Sphyrna mokarran) swimming over sandy seabed, South Bimini, Bahamas. The Bahamas National Shark Sanctuary, West Atlantic Ocean
Hoffmanns Two-toed sloth (Choloepus hoffmanni) mother and baby, aged 2 months, in tree, Costa Rica. Rescued and released by Aviarios Sloth Sanctuary
Proboscis monkey (Nasalis larvatus) mother suckling infant, Sabah, Malaysia
Harpy eagle (Harpia harpyja) close up head portrait, captive
Lammergeier / Bearded vulture (Gypaetus barbatus) flying, Leukerbad, Valais, Switzerland, February. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Song Thrush (Turdus philomelos), Bayern, Germany. June. June
Pilot whales (Globicephala macrorhynchus) just below surface, Tenerife, Canary Islands, Atlantic Ocean
Short-finned pilot whales (Globicephala macrorhynchus) two just below surface, Los Gigantes, South Tenerife, Canary Islands, Atlantic Ocean, July
Sheet ice crystals viewed by polarised light
Coral (Euphyllia ancora) polyps brown and green. Lembeh Strait, North Sulawesi, Indonesia
Red spider mite (Tetranychus urticae), Andalusia, Spain, June
Ghost glass frog (Sachatamia ilex), Costa Rica
Jaguar (Panthera onca) female yawning, native to Southern and Central America, captive
Horsesgrazing on Bulbarrow Hill at dawn, Dorset, England, UK, July 2014
Jungle cat (Felis chaus), female in search of prey. Velavadar National Park, Gujarat, India
Magnificent hummingbird (Eugenes fulgens) female, Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Fiery throated hummingbird (Panterpe insignis) an endemic bird species. Talamanca Range, Talamanca Range-La Amistad Reserves / La Amistad National Park UNESCO Natural World Heritage Site, Costa Rica
Northern pitcher plant (Sarracenia purpurea) photographed on Borgles Island, Nova Scotia, Canada, September
Red oak (Quercus rubra) low angle view of trees, New Brunswick, Canada, May
Alpine ibex (Capra ibex) close-up portrait. Gran Paradiso National Park, the Alps, Italy. January. Highly commended in the Portfolio category of the Terre Sauvage Nature Images Awards 2017
Magpie inkcap (Coprinopsis picacea) uncommon inkcap that usually grows singularly often under beech trees, Bedfordshire, England, UK, October. Focus stacked image
Reed warbler (Acrocephalus scirpaceus) with insect larvae in beak, Greylake RSPB Reserve, Somerset Levels, England, UK, June
Red water caused by tannins in peat bog flowing out of the Etang de la Gruere, peat bog, Rhine river catchment, Canton of Jura, Switzerland. November 2013. Photographed for the Freshwater Project
Giant panda (Ailuropoda melanoleuca) female, Huan Huan, holding baby, aged two months, Beauval Zoo, France, October 2017
Black bears (Ursus americanus) standing on back legs, fighting, Minnesota, USA, June
American flamingo (Phoenicopterus ruber) pair in courtship, group of three feeding, Punta Cormorant, Floreana Island, Galapagos
Waved albatross (Phoebastria irrorata) group of three on nest, Punta Suarez, Espanola Island, Galapagos
Galapagos penguin (Spheniscus mendiculus) incubating egg, Puerto Pajas, Isabela Island, Galapagos
Barn swallow (Hirundo rustica) feeding young, Monmouthshire, Wales, UK, July
