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Suspension bridge across the river, entrance to Gunung Mulu National Park UNESCO Natural World Heritage Site, Malaysian Borneo
Landscape modified by the erosion process. Goreme National Park and the Rock Sites of Cappadocia UNESCO World Heritage Site. Turkey. December 2006
Common redpoll (Acanthis flammea), flock feeding, Finland, January
Southern Rockhopper penguin (Eudyptes chrysocome) colony, Kidney Island, Falkland Islands, October. Vulnerable species
Silver Drummer (Kyphosus sydneyanus) fish off the coast of Neds Beach, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site, New South Wales, Australia, October 2012
Gentoo penguin (Pygoscelis papua) walking back to the colony, Sea Lion Island, Falkland Islands, October
Mute swan (Cygnus olor) cygnets with head submerged while feeding underwater, with mother watching and helping, Derbyshire, England, UK. June
Willow (Salix) plantation, Cuenca, Castilla-La Mancha, Spain, December
Fallow deer (Dama dama) at night, Gyulaj, Hungary Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Ordesa y Monte Perdido National Park, Huesca, Aragon, Spain, July 2016
Bornean orangutan (Pongo pygmaeus) male, Gunung Palung National Park, Borneo
Eurasian griffon vulture (Gyps fulvus) in flight, Cevennes, France, March 2016
Wild boar (Sus scrofa) running across snow covered field, Vosges, France, January
Leaf mimic bush cricket or katydid (unknown species, family Tettigoniidae) in the middle of moulting its skin. Moulting from one sub-adult stage (instar) to another is known as ecdysis
Giant Manta Ray (Manta birostris), with Remoras (Echeneididae) Socorro Island, Revillagigedo Archipelago Biosphere Reserve (Socorro Islands), Pacific Ocean, Western Mexico, March
White fronted capuchin monkey (Cebus capucinus) feeding, March, Costa Rica
Madagascar / Malagasy kingfisher (Alcedo vintsioides) perching, Anjajavy Private Reserve, north west Madagascar
Afrian lioness (Panthera leo) playing with her juvenile cub aged 2 years in Duba Plains concession. Okavango delta, Botswana
Wicken Fen wetlands with windmill water pump and Phragmites reeds, Cambridgeshire, England, May
River through Virgin Komi Forests UNESCO World Heritage site, the largest virgin forests in Europe. Ural Mountains, Komi Republic, Russia. August 2016
Meerkats (Suricata suricatta), Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
White rhino (Ceratotherium simum) at waterhole at night, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Young ground squirrels (Xerus inauris) fighting, Kgalagadi Transfrontier Park, Northern Cape, South Africa, January
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Eastern Ring-tailed Vontsira (Galidia elegans elegans) (endemic family Eupleridae) foraging on forest floor. Marojejy National Park, north east Madagascar
Adult King Vulture (Sarcoramphus papa) Laguna del Lagarto, Boca Tapada, Caribbean slope, Costa Rica, Central America
Female Red-eyed Tree Frog (Agalychnis callidryas) - Caribbean slope race (blue flanks) - obscured in rainforest foliage. Mid-altitude rainforest near Aranal, central Costa Rica
White-headed capuchin (Cebus capucinus) walking on giant palm-tree, Costa Rica
Common dolphin (Delphinus delphis) close up of dolphin surfing at speed, Hebrides, Scotland, UK
Peringueys desert adder (Bitis peringueyi) camouflaged in desert sand, Namibia
Kori bustard (Ardeotis kori) male courtship display, Kgalagadi Transfrontier Park, Northern Cape, South Africa
Bluethroat (Luscinia svecica) singing on a branch Vendee, France, May
Misty morning view of Exmoor countryside from Winsford Hill, Exmoor National Park, Somerset, UK. October 2016
Freshwater isopods (Acanthogammarus victorii) feeding on seabed, Lake Baikal, Siberia, Russia
Young Stone sculpin (Paracottus knerii), on endemic sponge (Lubomirskia baicalensis), Lake Baikal, Siberia, Russia
Clarion Lizard (Urosaurus clarionensis) displaying, IUCN Vulnerable, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Seychelles Sunbird (Cinnyris dussumieri), adult male singing, Praslin Island, Republic of Seychelles
Indian wild ass (Equus hemionus khur), group standing together, Little Rann of Kutch, Gujarat, India
Sea cucumber / Sea apple (Pseudocolochirus violaceus) Rinca, Komodo National Park, Indonesia
Triglav gentian (Gentiana terglouensis) Passo di Falzarego, near Cortina, Dolomites, Veneto, Italy. July
Tropical Flounder (Bothus mancus), Socorro Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands), Pacific Ocean
Pristine tropical rainforest and granite cliffs in mist, Morne Seychelles National Park, Mahe Island, Republic of Seychelles
Stripeless day gecko (Phelsuma astriata), camouflaged between leaves, Praslin Island, Republic of Seychelles
Galapagos Shark (Charcharhinus galapagensis), Roca Partida islet, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Starling (Sturnus vulgaris), eating fruit of Japanese Persimmon (Diospyros kaki), Provence, France
Galapagos giant tortoise (Geochelone nigra porteri), Cerro El Chato, Santa Cruz Island, Galapagos Islands
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Common midwife toad (Alytes obstetricans), male toad carrying eggs, Vaucluse, France
