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Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Mirrored portrait of a Weedy scorpionfish (Rhinopias frondosa). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea. Digitally manipulated (the face is reflected down mid-line)
Imbabura Tree Frogs (Boana picturatus) perched on a branch. Bilsa, Esmeraldas, Ecuador
Coquerels sifaka (Propithecus coquereli) Palmarium, Atsinanana, Madagascar
Iguazu Falls, Iguacu National Park, Brazil, Argentina November 2016. Photographed for The Freshwater Project
Three Sea otters (Enhydra lutris) resting on ice, Alaska, June
Bluebell (Hyacinthoides non-scriptus) woodland abstract, Norfolk, May
Wood anemone flower (Anemone nemorosa), RHS Rosemoor, near Great Torrington, Devon, UK. April 2017
Elephants (Loxodonta africana) at waterhole drinking at night. One spraying water from trunk, Zimanga Private Game Reserve, KwaZulu-Natal, South Africa
Spinner dolphin (Stenella longirostris) engaged in spinning manoeuvre, Sri Lanka. Composite sequence of images
Reflections of clouds and landscape in Loch Arkaig, Glen Dessary, Lochaber, Scotland, UK, September 2016
Waterlilies (Nymphaea alba) underwater in lake. Alps, Ain, France
Spinner dolphin (Stenella longirostris) juvenile surfacing next to its mother, Sri Lanka
Humpback whale (Megaptera novaeangliae) rear view of males escort, with adult female and male calf in the background. Vava'u, Tonga, South Pacific
Indo-Pacific bottlenose dolphin (Tursiops aduncus) leaping out of the surf, South Africa, Indian Ocean
Stac Pollaidh and Loch Lurgainn with Achiltibuie and Summer Isles beyond, looking west from Cul Beag, Coigach, Wester Ross, Scotland, UK, December 2016
Yellow pygmy goby (Lubricogobius exiguus) portrait, Kashiwajima, Japan
Large aggregation of Sperm whales (Physeter macrocephalus) engaged in social activity. Note the abundance of sloughed off skin in the water, resulting from tactile contact among the whales
Colony of Common Mussels (Mytilus edulis) growing on striated rock formation exposed at low tide. Cornwall, England, UK. Highly commended in the Coast
Swallowtail butterfly (Papilio machaon) flying to red campion with proboscis extended, Finland, June
Fragments of elkhorn coral (Acropora palmata) and staghorn coral: Acropora cervicornisgrowing, hung on coral propagation tree, as part of a coral conservation nursery project. East End, Grand Cayman
Pricklepear (Opuntia sp. ) growing on coastline, Socorro Island, Revillagigedo ArchipelagoPricklepear (Opuntia sp.) growing on coastline, Socorro Island, Revillagigedo Archipelago National Park (Socorro Islands), Pacific Ocean, Western Mexico, November
Aerial view of Espiritu Santo Island National Park, Sea of Cortez (Gulf of California), Mexico, June, Honorary Mention in the third national contest Visions of our Nature organised by Mexicos
Green Turtle (Chelonia mydas) trapped in fishing net, Sea of Cortez (Gulf of California), Mexico, May, Third Place in the third National Contest Visions of our nature organised by Mexicos National
Black-browed Albatross (Thalassarche melanophris) resting close up of eye. West Point Island, Falkland Islands, October
Pinnacles, also known as fairy chimneys or hoodoo, Love Valley. Goreme National Park and the Rock Sites of Cappadocia UNESCO World Heritage Site. Turkey. December 2006
African elephant mother and young calf (Loxodonta africana), Duba Plains, Okavango Delta, Botswana, Southern Africa
Common redpoll (Acanthis flammea) two birds perched, Finland, January
Northern hawk-owl (Surnia ulula) perched, Finland, January
Landscape of a fen at dawn withCanada goose (Branta canadensis). Oisterwijkse Bossen en Vennen Nature reserve, Oisterwijk, The Netherlands. April
Magellanic penguin (Spheniscus magellanicus) calling at the entrance of burrow, courtship display, Carcass Island, Falkland Islands, October
Whooper swan (Cygnus cygnus) on river in winter, Finland, January
Upland goose (Chloephaga picta) goslings, Sea Lion Island, Falkland Islands, October
King penguin (Aptenodytes patagonicus) chick creche, Volunteer Point, East Falkland, Falkland Islands, October
Striated caracara (Phalcoboenus australis) group with one landing from above with only wings and tail visible. Carcass Island, Falkland Islands, October
King penguin (Aptenodytes patagonicus) colony, Volunteer Point, East Falkland, Falkland Islands, October
Aerial view of fish farm, La Guittiere Marsh, South Vendee, France, July 2017
Leaves of a Common aspen tree (Populus tremula) on ground in autumn, Aragon, Spain, November
Footpath sign in Norfolk countryside with Blackthorn hedgerow (Prunus spinosa) in flower. Norfolk, England, UK. April
Eurasian blackcaps (Sylvia atricapilla) caught on limed stick, Cyprus. Finalist in The Wildlife Photojournalist Award: Single Image Category of the Wildlife Photographer of the Year Awards (WPOY)
African lion (Panthera leo) at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Zebra at waterhole at night, Mkuze, South Africa Third place in the Nature Portfolio category of the World Press Photo Awards 2017
Northern Lights (Aurora Borealis) over Hornafjordur, eastern Iceland, February 2016
Polar bear (Ursus maritimus) on an iceberg, Baffin Bay, Canada. September
Red leaf monkey (Presbytis rubicunda) female feeding in Strangler fig tree (Ficus dubia) Gunung Palung National Park, Kalimantan, Borneo, Indonesia
European robin (Erithacus rubecula), Vosges, France, December
Hawksbill turtle (Eretmochelys imbricata) cruising along the drop off of a coral reef. Bloody Bay Wall, Little Cayman, Cayman Islands. British West Indies. Caribbean Sea
Blue morpho (Morpho helenor) butterflies feeding on mango fruit, butterfly observatory, La Paz Waterfall Gardens, Costa Rica
