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White prominent moth (Leucodonta bicoloria) County Kerry, Ireland
Wren (Troglodytes troglodytes) on frost covered fern, Dunham Massey, Cheshire, UK. Winner of the Habitat Category of the British Wildlife Photographer of the Year Awards (BWPA) 2017
Brown bear (Ursus arctos arctos) female and two cubs, aged 1 year, emerging from den in winter, Bavarian Forest, Germany, captive, January
Reflections in Llyn Gwynant in the Glaslyn valley, looking west with Yr Aran mountain in the background, Snowdonia National Park, North Wales, UK, June
Accumulation of land snails (Euryptyxis labiosa), escaping the desert heat by resting on a shrubs bark, Ayn Sahnawt, Sultanate of Oman, February
Frill-necked lizard (Chlamydosaurus kingii), with collar spread in threat display. Alice Springs, Northern territory, Australia
Eurasian beaver (Castor fiber) sits up, captive, Devon, UK, May
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, lake Victoria, Uganda
Oak tree (Quercus robur) silhouetted against night sky with stars, Brecon Beacons National Park, Wales, UK. December
Humpback whale calf (Megaptera novaeangliae) male resting beneath his resting mother. Vava u, Tonga, South Pacific
Army ant (Eciton burchellii) major soldier, Parque da Onca Parda Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Male Golden Pheasant (Chrysolophus pictus) standing on grass in display plumage. Kew Gardens, England, UK. March
Crocodile monitor (Varanus salvadorii) close up eye, captive, occurs in New Guinea
Flowers (Trollius sp) Balang Mountain, Wolong National Nature Reserve, Sichuan Giant Panda Sanctuaries - Wolong, Mt Siguniang and Jiajin Mountains UNESCO World Heritage Site, Sichuan Province, China
Hammerhead worms (Bipalium sp) on leaf, Sarawak. Borneo
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Long-tailed duck (Clangula hyemalis), males in flight, Finland, April
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, August
Common swift (Apus apus) group flying overhead, Bradford-on-Avon, Wiltshire, UK, May
Norfolk Island pine trees (Araucaria heterophylla) near lagoon with Mount Lidgbird and Mount Gower in the background, Lord Howe island, Lord Howe Island Group UNESCO Natural World Heritage Site
Saffron crocuses (Crocus sativus), cultivated for saffron, Lleida, Catalonia, Spain, November
Common European toad tadpoles (Bufo bufo), sequence showing development of limbs. Derbyshire, UK. August. Digital composite
White-throated magpie-jay (Calocitta formosa) Nicoya peninsula, Costa Rica. February
Adult Aye-aye (Daubentonia madagascariensis) active in forest canopy at night. Dry deciduous forest near Andranotsimaty. Daraina, northern Madagascar. Critically Endangered
Close up view of Honey Bee comb showing larvae in cells Norfolk, England, June 2017
Pairs of Scarlet Macaws (Ara macao) in flight. Osa Peninsula (near Corcovado National Park), Costa Rica, Central America
Canebreak treefrog (Aplastodiscus sp) Guainumbi Private Reserve, Sao Paulo, Atlantic Forest South-East Reserves, UNESCO World Heritage Site, Brazil
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June
Peony (Paeonia officinalis) Mt Vettore, Umbria, Italy, June
Commensal emperor shrimp (Periclimenes imperator) hitching a ride on a Nudibranch (Risbecia tryoni). Lembeh Strait, North Sulawesi, Indonesia
Clarion Island Whip Snake (Masticophis anthonyi), IUCN Critically Endangered, Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage
Palm spider (Nephila inaurata) in its web, female, Praslin Island, Republic of Seychelles
Grey wolf (Canis lupus), adult with cub, captive, USA
Grass snake(Natrix natrix) coiled round eggs, Alsace, France
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Scarlet ibis (Eudocimus ruber), in flight, Coro, Venezuela
Nurse shark (Ginglymostoma cirratum) portrait, resting on the sand in shallow water. Its barbels are clearly visible on its top lip. South Bimini, Bahamas. The Bahamas National Shark Sanctuary
Nudibranchs (Nembrotha chamberlaini) feeding on Tunicates (sea squirt) on a coral reef. Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Southern Viscacha (Lagidium viscacia) at rest, Isla Incahuasi in the Salar de Uyuni, altiplano, Bolivia
Waterwheel plant (Aldrovanda vesiculosa) a carnivorous aquatic plant with Mosquito larva
Millstones made from millstone grit beneath Stannage Edge, Hathersage, Derbyshire, November 2016
Emerald toucanet (Aulacorhynchus prasinus) Rio Platano Biosphere Reserve and UNESCO World Heritage Site, La Mosquitia, Honduras
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June