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Ripples of sand on a shallow seabed. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Humpback whale (Megaptera novaeangliae) just under surface, off Shetland, Scotland, UK. First ever underwater images of humpbacks in British waters
Kelp forest (Laminaria digitata) with small fish, Shetland, Scotland, UK, July
RF - St Michaels Mount and old causeway at sunrise, Marazion, Cornwall, UK. October 2015
European otter (Lutra lutra) on log over water, captive, the Netherlands
Young Platypus (Ornithorhynchus anatinus) is released onto a log in McMahons Creek, Yarra Ranges, Victoria, Australia. Photographed under controlled conditions. September
Shingle beach and chalk cliffs along the North Sea coast at Yport, Normandy, France, June 2018
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed. These octopus regularly carry pieces of shell so they can use them as protective
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Fin whale (Balaenoptera physalus) near surface. Pico Island, Azores, Portugal. May
Orcas / killer whales (Orcinus orca) swimming in open water, Three Kings Islands, New Zealand. Pacific Ocean
Florida manatee (Trichechus manatus latirostris) swimming into a fresh water spring, Three Sisters Spring, Crystal River, Florida, USA
RF - Mantis shrimp (Odontodactylus scyllarus) on walk about on coral reef. Puerto Galera, Philippines
Nudibranch (Phyllidia marindica), Lembeh Strait, North Sulawesi, Indonesia. December
Underwater view of Waterlilies (Nymphaea alba) in a lake. Alps, Ain, France, June
Spekes Mill Mouth waterfall near Hartland Quay, Devon, UK, March
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Rainbow river, Rainbow Springs State Park, Florida, USA. January. Photographed for The Freshwater Project
Sunrays in Cenote Jardin del Eden, Quintana Roo, Yucatan Peninsula, Mexico, May 2016. Photographed for The Freshwater Project
Swallow-tailed gull (Creagrus furcatus) Plazas Island, Galapagos
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise
Head close up of Black labrador retriever dog (field type) swimming in pond with autumn reflections, Rhode Island, USA
Robin (Erithacus rubecula) at RSPB Leighton Moss Bird Reserve, Silverdale, Lancashire, England, UK. February 2016
Landscape of Zanskar River, a tributary of the Indus, borders part of Hemis national park, habitat of the snow leopard (Panthera uncia), Hemis National Park, Ladakh, India
Yellow bellied toad (Bombina variegata) showing underside, Kirchheimbolanden, Germany. Meetyourneighbours.net project
Moose (Alces alces) bull swimming in water, Baxter State Park, Maine, USA
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Knoydart and Loch Nevis from Sgurr Cos na Breachd-laoidh with sun shining through cloud, Lochaber, Scotland, UK, September 2016
Oldshoremore Beach and dunes in evening light, Kinlochbervie, Sutherland, Scotland, UK, April 2014
Warebeth Beach at dawn with view to Hoy, Orkney, Scotland, UK, November 2014
Black-throated Diver (Gavia arctica) Rokua, Finland, May
Llyn Cregennen and view toward Cadair Idris, late evening lightr, near Dolgellau, Snowdonia National Park, Wales, UK, September 2013
Rough seas during Storm Ruth, Ile d Ouessant, Armorique Regional Park. Iles du Ponant, Finistere, Brittany, France, Iroise Sea. 8th February 2014
Pine marten (Martes martes) reflected in water, Ardnamurchan Peninsula, west coast of Scotland, UK. Highly commended in the Mammals category of the British Wildlife Photography Awards (BWPA)
RF- Goldfinch (Carduelis carduelis) reflected in pool, Worcestershire. November
Nudibranchs (Doto greenamyeri) newly described species on feather hydroids. Seraya Bay, north coast of Bali, Indonesia. Java Sea
School of Silversides (Atherinomorus lacunosus) mass below a jetty, creating a false ceiling of fish. Berenice Jetty, Aqaba, Jordan. Gulf of Aqaba, Jordan
Portrait of Bigfin squid (Sepioteuthis lessoniana) hovering in open water above a coral reef at night. Dampier Strait, Raja Ampat, West Papua, Indonesia. Tropical West Pacific Ocean
Underwater view of pebbles near the shore of Lake Baikal Lake Baikal UNESCO World Heritage Site, Siberia, Russia. May 2013. Photographed for The Freshwater Project
Pied avocet (Recurvirostra avosetta) feeding in shallow water at dawn, Etosha National Park, Namibia
Great crested grebe (Podiceps cristatus) carrying chick on back, The Netherlands, May
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Red sea fan (Melithaea sp.) is surrounded by Glassfish ( Apogon sp.) on a coral reef. Daram Islands, Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Stac Pollaidh, Cul Mor and Suilven, view north from Sgorr Tuath, Coigach and Assynt, Scotland, UK, March 2017
Portrait of Sally lightfoot crab (Grapsus grapsus) on a beach. San Cristobal Island, Galapagos Islands, Ecuador. Tropical East Pacific Ocean
Feigh Burn River, Dunseverick Falls flowing into the sea, County Antrim, Northern Ireland. November 2020
Portuguese man o war (Physalia physalis) Tenerife, Canary Islands
