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Larval Wonderpus octopus (Wunderpus photogenicus) drifting in the open ocean at night off Anilao, Philippines
RF Humpback whale (Megaptera novaeangliae) aerial view. Baja California, Mexico
Pygmy blue whale (Balaenoptera musculus brevicauda) subspecies of blue whale, diving downwards, Mirissa, Sri Lanka, Indian Ocean. Endangered species
Firefly squid (Watasenia scintillans) emitting light from photophores, Toyama Bay, Japan. This form of bioluminescence is used as counter illumination camouflage
Tasseled wobbegong shark (Eucrossorhinus dasypogon) yawning. Citrus Ridge, Yanggefo Island, Gam Island, Raja Ampat, West Papua, Indonesia
Gorgonian sea fans (Acalycigorgia sp) on coral reef at Raja Ampat, West Papua, Indonesia
Air filled bladders of Giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean
Red tailed boa constrictor (Boa constrictor) on fallen tree over water, Yasuni National Park, Orellana, Ecuador
Red squirrel (Sciurus vulgaris) at woodland pool, feeding on nut, Scotland, UK, November. Highly commended, Animal Portraits category, British Wildlife Photography Awards (BWPA) competition 2012
River Brue at dawn near Glastonbury on the Somerset Levels, Somerset, UK, November 2010
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise with bivalve shell
Underwater shot of aquatic plants in an oxbow lake of the River Aare, Haftli nature Reserve, Buren an der Aare, Switzerland
Bioluminescent Caribbean reef squid (Sepioteuthis sepioidea), St. Vincent, Eastern Caribbean
Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata)
Yellow tropical reef fish composite image on black background, Falcula butterflyfish (Chaetodon falcula), Yellow boxfish (Ostracion cubicus), Golden butterflyfish (Chaetodon semilarvatus)
Giant Pacific octopus (Enteroctopus dofleini) experiencing freedom after release from captivity, Vancouver Island, British Columbia, Canada, Pacific Ocean
Female Peacock-mantis shrimp (Odontodactylus scyllarus) carrying her eggs, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Aggregation of thousands of moulting Spider crabs (Leptomithrax gaimardii), South Australia Basin, Australia. Pacific Ocean
Compass jellyfish (Chrysaora hysoscella) with sunburst close to the surface, Cornwall, UK, .English Channel
Guineafowl puffer fish (Arothron meleagris) inflated, displaying defensive behaviour over coral reef, Yap, Indo Pacific, Micronesia
Fafiao Waterfalls, Peneda Geres National Park, Portugal, February 2009
Great crested grebe (Podiceps cristatus) adult with young on the back in the first morning light Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. May
Aerial view of southern tip of Dominica, Scotts Head. It divides Atlantic from Caribbean sea. Dominica, West Indies. October 2020
Blue dragon seaslug (Glaucus atlanticus) with Blue button hydroid colony (Porpita porpita) on which it feeds. Tenerife, Canary Islands. October
Achnambeithach Cottage on the shores of Loch Achtriochtan at the foot of Aonach Dubh, Glencoe, Lochaber, Highland, Scottish Highlands, Scotland, UK. June 2018
Aerial view of Red soda in Lake Magadi, Rift Valley, Kenya. October 2019
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) shoal swimming over Turtlegrass (Halophila stipulacea) seagrass bed. Marsa Alam, Egypt
Short-snouted seahorse (Hippocampus hippocampus) male, Ponza Island, Italy, Tyrrhenian Sea, Mediterranean Sea
Yellowstone River. Yellowstone National Park, Wyoming, USA. September
Slime mould (Physarum album), dew droplets on two sporangia, close-up. Hertfordshire, England, UK. November. Focus stacked image
Spiny dogfish (Squalus acanthias) portrait. Rhode Island, New England, USA. August
Salmon shark (Lamna ditropis). Port Fidalgo, Alaska, USA. July
Puffins (Fratercula arctica) swimming underwater. Puffins spend most of their lives at sea and are excellent underwater swimmers, which is how they catch small fish, their main food
Leafy seadragon (Phycodurus eques) male carrying eggs, swims over seagrass meadow. Wool Bay, Edithburgh, South Australia. Gulf of St Vincent
RF - American crocodile (Crocodylus acutus) swims through sunrays. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
Diver approaches a large colony of Elkhorn coral (Acropora palmata) growing in shallow water on a coral reef. Wreck of the Glamis, East End, Grand Cayman, Cayman Islands, British West Indies
Green sea turtle (Chelonia mydas) swimming near surface in halo of light. The Bahamas
Deepsea jellyfish (Periphylla periphylla) juvenile, , Trondheims fjord, Norway, Atlantic ocean
Fire salamander (Salamandra salamandra) in forest habitat, Hallerbos, Belgium. May
Mandarin duck (Aix galericulata) male female pair, Yuyuantan Park, Beijing, China
Split level photo of an American crocodile (Crocodylus acutus) beneath red mangrove trees (Rhizophora mangle) above a bed of seagrass (turtlegrass: Thalassia testudinum)
Giant hogfish (Bodianus macrognathos) male yawning at a cleaning station run by juvenile Spanish hogfish (Bodianus rufus). Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
A yellow tube sponge (Aplysina fistularis) growing on a Caribbean coral reef, withsun burst above. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
Sea elephant (Cardiopoda placenta), marine pelagic gastropod mollusc in the family Pterotracheidae. Deep sea heteropod. Atlantic Ocean, close to Cape Verde. Controlled conditions
Amphipod (Acanthogammarus lappaceus) Lake Baikal, Siberia, Russia. Endemic to Lake Baikal
Lake baikal sponge and a mixture of freshwater snails Lake Baikal, Siberia, Russia. August
Amphipod (Acanthogammarus victorii), Lake Baikal, Russia. June. Endemic to Lake Baikal
