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Pair of Caribbean reef sharks (Carcharhinus perezi) swim over a coral reef. Jardines de la Reina, Gardens of the Queen National Park, Cuba. Caribbean Sea
White chinned petrel (Procellaria aequinoctialis) in flight low over the water off Kaikoura, Canterbury, New Zealand
Fallow deer (Cervus dama) stags at sunset, Holkham Park, North Norfolk, UK, January
Green turtles (Chelonia mydas) swimming with angelfish, Punta Vicente Roca, Isabela Island, Galapagos
Great blue heron (Ardea herodias) taking off, with coastal landscape, Las Bachas, Santa Cruz Island, Galapagos
Starr Puffer (Arothron stellatus), juvenile swimming. Lembeh Strait, North Sulawesi, Indonesia
Reef manta rays (Manta alfredi) filter feeding in atoll passes and lagoons Hanifaru Bay, Raa Atoll, Maldives
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
Jellyfish (Diplulmaris antarctica) Cuverville, Antarctic Peninsula, Antarctica
Lined seahorses (Hippocampus erectus) amongst corals, The Bahamas
Sanderling (Calidris alba) flock foraging at sunset, Britanny, France, October
American lobster (Homarus americana) in eelgrass (Zostera marina). Nova Scotia, Canada. July
Village Bay on St Kilda and the old house that formed the village, Scotland, UK, May
Three North American / Canadian Otters (Lutra canadensis) lying on each other by water. Captive, UK
Carp (Cyprinus carpio) in oxbow lake of the Aare River, Switzerland
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Grey whale (Eschrichtius robustus) tail, San Ignacio Lagoon, El Vizcaino Biosphere Reserve, Baja California, Mexico, March
Coal tit (Periparus ater) on mossy log with reflection, England, UK. January
Ewens Ponds, spring fed limestone ponds at the Ewens Ponds Conservation Park, South Australia, Australia March 2015. Photographed for The Freshwater Project
Brown trout (Salmo trutta) in shallow water migrating upstream, Bornholm, Denmark, November 2008
Common dolphins (Delphinus delphis) surfacing, Fisheye lens. Pico, Azores, Portugal, June 2009
Sgwd Isaf Clun-gwyn waterfall and rapids. Ystradfellte, Brecon Beacons National Park, Wales, November 2011
RF- Mute swan (Cygnus olor) on water in winter dawn mist. Loch Insh, Cairngorms National Park, Highlands, Scotland UK, December
View from A Mhaighdean overlooking Fionn Loch. Highlands, Highlands of Scotland, UK, January 2016
Kamchatcka brown bear (Ursus arctos beringianus) with face underwater whilst fishing for Sockeye salmon in the Ozernaya River, Kuril Lake, South Kamtchatka Sanctuary, Russia, August
Reflections in Llyn Gwynant on a misty morning, Glaslyn valley, with Yr Aran mountain in the background, Snowdonia National Park, North Wales, UK, November 2017
Red squirrel (Sciurus vulgaris) having a drink, Black Isle, Scotland, UK. October
Kingfisher (Alcedo atthis) taking off from water with caught fish. Worcestershire, UK, March
Great crested grebe (Podiceps cristatus) pair performing the weed dance, a spectacular part of their elaborate courtship ritual, Derbyshire, UK, March
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Corys shearwaters (Calonectris diomedea) diving among a mass of shoaling fish to feed, along with Atlantic spotted dolphins (Stenella frontalis), Formigas Islet dive site, Azores, Portugal
Adelie penguin (Pygoscelis adeliae) diving near ice flow, Antarctica. Small reproduction only
King Eider (Somateria spectabilis) flying low over water. Floe edge, Arctic Bay, Nunavut, Canada, April
Portrait of a Green sea turtle (Chelonia mydas) in morning light. Sipadan Island, Sabah, Borneo, Malaysia. Celebes Sea
Green Turtle, (Chelonia mydas), Swimming over volcanic sandy bottom, Armenime cove, South Tenerife coast, Canary Islands, Spain, Atlantic Ocean, May 2011
Great crested grebes (Podiceps cristatus) performing courtship ritual at dawn, Cheshire, UK April
Man-of-war fish (Nomeus gronovi) living within deadly tentacles of Portuguese man o war (Physalia physalis), to whose toxin it is ten times more resistant than other fish, at sea surface. Dominica
Mimika bobtail squid (Euprymna morsei), Hokkaido, Japan, Pacific Ocean
Australian sea (Neophoca cinerea) juvenile playing in the sand, Carnac Island, Western Australia
Pelagic thresher shark (Alopias pelagicus) swimming in open ocean, Malapascua Island, Philippines, Pacific Ocean
Common loon (Gavia immer), in breeding plumage, running across water to take flight, Michigan, USA. June
Giant pacific octopus (Enteroctopus dofleini) swimming freely after release from captivity, Vancouver Island, British Columbia, Canada, Pacific Ocean
Spotted eagle rays (Aetobatus narinari) swimming, Hawaii, USA
Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Bottlenose dolphins (Tursiops truncatus) at the surface at sunrise. Kwazulu-Natal, South Africa
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
