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African pompano (Alectis ciliaris) juvenile with long trailing dorsal and anal fins, swimming in mid-water at night. In-camera double exposure
Moon jelly (Aurelia aurita) drifts over a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
RF - African elephants (Loxodonta africana) in water, trunks touching, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean
Dugong (Dugong dugon) male and juvenile Golden trevally (Gnathanodon speciosus) swimming over Seagrass (Halophila stipulacea) meadow. Marsa Nabaa, Marsa Alam, Egypt
Canada Goose (Branta canadensis) in flight. Gilbert Water Ranch, Arizona, USA. January
Western lowland gorilla (Gorilla gorilla gorilla) male aged 12 years standing up at edge of Mpassa river. Captive bred in UK and released into wild along with his father in 2013
Malabar Torrent toad (Ghatophryne ornata) in habitat, Amboli, Maharashtra, India. Western Ghats. Endangered and endemic to Western Ghats
Orca (Orcinus orca) with South American sealion (Otaria flavescens) in mouth, beaching itself on shore. Punta Norte, Valdez Peninsula, Argentina. April
Mandarin duck (Aix galericulata) pair mating on lake. Shanghai Zoo, China
Bubble eye goldfish (Carassius auratus) with upward pointing eyes and two large fluid-filled sacs.. In captivity, Shanghai Zoo, Shanghai, China
Chinese grass carp / White amur (Ctenopharyngodon idella), farmed for food in China. Captive specimen, China
Gayal / Mithun (Bos frontalis) on river bank. Arunachal Pradesh, North East India. November 2014
Transparent black ice with cracks on Lake Baikal, aerial shot. Photographed for The Freshwater Project extended. Near Khuzhir, Olchon Island, Irkutsk Oblast and Buryat Republic, Siberia, Russia
Aardvark (Orycteropus afer) drinking at night, Zimanga private game reserve, KwaZulu-Natal, South Africa. Medium repro only
RF - Green sea turtle (Chelonia mydas) Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean. (This image may be licensed either as rights managed or royalty free.)
Portrait of a Giant frogfish (Antennarius commersoni) on a large Yellow elephant ear sponge (Ianthella basta). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Blue whale (Balaenoptera musculus) swims beneath the surface of the ocean. Indian Ocean, off Sri Lanka
Portrait of a large female Reef manta ray (Mobula alfredi) swimming over a coral reef. Misool, Raja Ampat, West Papua, Indonesia. Ceram Sea. Tropical West Pacific Ocean
Free swimming mimic octopus (Thaumoctopus mimicus). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Beaver Bridge over River Conwy, near Betws-y-Coed, Snowdonia National Park, North Wales, UK, September 2018
Autumnal boreal forest with lake, Silver Trail, near Mayo, Yukon Territories, Canada, September 2013
Beech trees (Fagus sylvaticus) and pond in autumn, Waggoners Wells, Surrey, England, UK, October
Blue-footed booby (Sula nebouxii) diving towards sea, folding wings. Northeast coast, Santiago Island, Galapagos. Sequence 1/2
Humpback whale (Megaptera novaeangliae) mother with young calf in tropical sheltered coastal waters Vava u, Tonga
Blue shark (Prionace glauca) with a pair of pilot fish (Naucrates ductor) off Halifax, Nova Scotia, Canada. July
Sugar kelp (Sacchinaria latissima) off Nova Scotia, Canada. September
Moorish idol (Zanclus cornutus) juvenile with transparent skin in the open ocean at night off Anilao, Philippines
Pelagic thresher shark (Alopias pelagicus) being cleaned by a bluestreak cleaner wrasse (Labroides dimidiatus) at Monad Shoal off Malapascua Island, Philippines
Wonderpus octopus (Wunderpus photogenicus) in its juvenile or larval stage. At this stage it is transparent and drifts in the open ocean as plankton. Anilao, Philippines
Ribbon eel ( Rhinomuraena quaesita) Lembeh Strait, North Sulawesi, Indonesia
Nudibranch (Chromodoris willani) with a scale worm. Lembeh Strait, North Sulawesi, Indonesia
Corkscrew or Long tentacle anemone (Macrodactyla doreensis). Lembeh Strait, North Sulawesi, Indonesia
Taylors pygmy leatherjacket (Brachaluteres taylori). Lembeh Strait, North Sulawesi, Indonesia
Water lilies (Nymphaea alba) lit by the suns rays, seen under water. Alps, Ain, France. June. Highly commended in the the Underwater World category
Kelp forest (Laminaria hyperborea) in the clear seas of Scotland, UK, September
Common buzzard (Buteo buteo) landing on prey, wings open, Scotland, UK, September 2016
Small lochan surrounded by flowering heather (Ericaceae sp) on the edge of the Caledonian pine forest, Rothiemurchus, Cairngorms NP, Scotland, UK, September 2011
Snares island crested penguin (Eudyptes robustus) colony on the coast, high angle view, Snares Island, New Zealand
Tiger (Panthera tigris tigris) dominant male tiger approaching Beja pani waterhole, Kanha National Park, Central India. Camera trap image
Puffins (Fratercula arctica) Shaint Isles, Scotland, UK, June
Common dolphin (Delphinus delphis) jumping out the water, Shetland, Scotland, UK, January
Big storm hitting cliffs, with waves breaking over the top, Shetland, Scotland, UK, July
Jellyfish (Pelagia noctiluca) amongst plankton, Shetland Isles, Scotland
Grey heron (Ardea cinerea) hunting for food amongst moss with loch and Celtic rainforest in back ground, Scotland, UK, October
Orca (Orcinus orca) surfacing and blowing, Shetland, Scotland, UK, August
Green sandpiper (Tringa ochropus) with small worm it has caught in shallow freshwater lake, Gloucestershire, UK, August