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Barbary macaque (Macaca sylvanus), two adults with baby, overlooking town and ships in sea at sunset. Gibraltar Nature Reserve, Gibraltar. August 2018
Bottlenose dolphins (Tursiops truncatus) at the surface at sunrise. Kwazulu-Natal, South Africa
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Aggregation of Golden jellyfish (Mastigias sp.) in a marine lake in Palau, the golden colour of this species comes from symbiotic algae in its tissues
Reflections in Llyn Gwynant in the Glaslyn valley, looking west with Yr Aran mountain in the background, Snowdonia National Park, North Wales, UK, June
A wide angle macro view of a Veined octopus ( Amphioctopus marginatus) stretching out from its protective den in an old clam shell. Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea
Clarks Grebe (Aechmophorus clarkii) with yawning chick riding on back, Bear River Refuge, Utah, July
Bog wetlands with Suilven mountain in the background at dawn, Assynt mountains, Highland, Scotland, UK, June 2011
Sunrise over coastal mudflats, Campfield Marsh RSPB reserve, Bowness, Solway Firth, Cumbria, UK, April 2011
Whooper swan (cygnus cygnus) silhouette at sunrise, Loch Insh, Cairngorms NP, Kincraig, Scotland, UK, March 2011
White-throated dipper (Cinclus cinclus) winter with river flooding, Kuusa, Laukaa, Keski-Suomi, Lansi- ja Sisa-Suomi / Central and Western Finland, Finland. January
High tide at Trebarwith Strand at sunset, North Cornwall, UK. May 2013
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Northern gannets (Morus bassanus) in flight in Force 8 gales above raging seas. Shetland Islands, Scotland, UK, September
Grey Seal (Halichoerus grypus) at rest on submerged rock, head and one flipper above water. Shetland Islands, Scotland, UK. July
Watermilfoil (Myriophyllum spicatum) with some stems supporting clusters of Zebra mussel (Dreissena polymorpha) Lake Neuchatel, Switzerland
Green turtle (Chelonia mydas) swimming near mangroves, Elizabeth Bay, Isabela Island, Galapagos
Mute swan (Cygnus olor) parents and cygnets on a misty lake at sunrise. London, UK. May
Ocean sunfish (Mola mola) off Halifax, Nova Scotia, Canada. July
Schools of Pacific creolefish (Cephalopholis colonus) and Panamic sergeant major (Abudefduf troschelii) swimming through sun beams inside an underwater cavern
Red mangrove (Rhizophora mangle) propagules / plantlets which become fully mature plants before dropping off the parent tree to drift away and establish a new tree. Bahamas
European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July
Larval Wonderpus octopus (Wunderpus photogenicus) drifting in the open ocean at night off Anilao, Philippines
RF Humpback whale (Megaptera novaeangliae) aerial view. Baja California, Mexico
Pygmy blue whale (Balaenoptera musculus brevicauda) subspecies of blue whale, diving downwards, Mirissa, Sri Lanka, Indian Ocean. Endangered species
Firefly squid (Watasenia scintillans) emitting light from photophores, Toyama Bay, Japan. This form of bioluminescence is used as counter illumination camouflage
Tasseled wobbegong shark (Eucrossorhinus dasypogon) yawning. Citrus Ridge, Yanggefo Island, Gam Island, Raja Ampat, West Papua, Indonesia
Gorgonian sea fans (Acalycigorgia sp) on coral reef at Raja Ampat, West Papua, Indonesia
Air filled bladders of Giant kelp (Macrocystis pyrifera). Santa Barbara Island, Channel Islands. Los Angeles, California, United States of America. North East Pacific Ocean
Red tailed boa constrictor (Boa constrictor) on fallen tree over water, Yasuni National Park, Orellana, Ecuador
Red squirrel (Sciurus vulgaris) at woodland pool, feeding on nut, Scotland, UK, November. Highly commended, Animal Portraits category, British Wildlife Photography Awards (BWPA) competition 2012
River Brue at dawn near Glastonbury on the Somerset Levels, Somerset, UK, November 2010
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise with bivalve shell
Underwater shot of aquatic plants in an oxbow lake of the River Aare, Haftli nature Reserve, Buren an der Aare, Switzerland
Bioluminescent Caribbean reef squid (Sepioteuthis sepioidea), St. Vincent, Eastern Caribbean
Blue tropical reef fish composite image on black background, Blue triggerfish (Pseudobalistes fuscus), Bicolour angelfish (Centropyge bicolor), Elongate surgeonfish (Acanthurus mata)
Yellow tropical reef fish composite image on black background, Falcula butterflyfish (Chaetodon falcula), Yellow boxfish (Ostracion cubicus), Golden butterflyfish (Chaetodon semilarvatus)
Giant Pacific octopus (Enteroctopus dofleini) experiencing freedom after release from captivity, Vancouver Island, British Columbia, Canada, Pacific Ocean
Female Peacock-mantis shrimp (Odontodactylus scyllarus) carrying her eggs, Raja Ampat, West Papua, Indonesia, Pacific Ocean
Aggregation of thousands of moulting Spider crabs (Leptomithrax gaimardii), South Australia Basin, Australia. Pacific Ocean
Compass jellyfish (Chrysaora hysoscella) with sunburst close to the surface, Cornwall, UK, .English Channel
Guineafowl puffer fish (Arothron meleagris) inflated, displaying defensive behaviour over coral reef, Yap, Indo Pacific, Micronesia
Fafiao Waterfalls, Peneda Geres National Park, Portugal, February 2009
Great crested grebe (Podiceps cristatus) adult with young on the back in the first morning light Valkenhorst Nature Reserve, Valkenswaard, The Netherlands. May
Aerial view of southern tip of Dominica, Scotts Head. It divides Atlantic from Caribbean sea. Dominica, West Indies. October 2020
Blue dragon seaslug (Glaucus atlanticus) with Blue button hydroid colony (Porpita porpita) on which it feeds. Tenerife, Canary Islands. October
Achnambeithach Cottage on the shores of Loch Achtriochtan at the foot of Aonach Dubh, Glencoe, Lochaber, Highland, Scottish Highlands, Scotland, UK. June 2018
