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Sally lightfoot crabs (Grapsus grapsus) on rock with waves, Cape Douglas, Fernandina Island, Galapagos
Backswimmer (Notonecta glauca), resting at the water surface, Europe, August, controlled conditions
Great diving beetle (Dytiscus marginalis) and Dragonfly nymph (Aeshnidae), Europe, August, controlled conditions
Great diving beetle (Dytiscus marginalis) male, Europe, August, controlled conditions
Pederson cleaner shrimp (Periclimenes pedersoni), Cienaga de Zapata National Park, Matanzas Province, Cuba
View of the Rio Celeste waterfall, tropical rainforest of Tenorio Volcano National Park, Costa Rica. The blue color arises due to a physical phenomenon known as Mie scattering triggered by
Unicorn leatherjacket filefish (Aluterus monoceros) at night, The Bahamas. February
Common octopus (Octopus vulgaris) on a coral reef, The Bahamas. August
Common octopus (Octopus vulgaris) on a coral reef in The Bahamas. August
Aerial view of tropical rainforest and meandering river, Salonga National Park, Democratic Republic of Congo May 2017
Tiger moray / Fangtooth Moray (Enchelycore anatina) and Black moray eel (Muraena augusti), South Tenerife, Canary Islands, Atlantic Ocean
Couple of Blue shark (Prionace glauca) and Pilot fish (Naucrates ductor), Pico Island, Azores, Portugal, Atlantic Ocean
Blue shark (Prionace glauca) with fishing hook caught in mouth and Pilot fish (Naucrates ductor), Pico Island, Azores, Portugal, Atlantic Ocean
Face portrait of a Great hammerhead shark (Sphyrna mokarran) off Bimini, Bahamas
Veined Octopus, Coconut Octopus (Amphioctopus marginatus, formerly Octopus marginatus), inhabiting discarded bivalve mollusc shells, which it collects and pulls around itself - defense, shelter
Giant tun (Tonna galea) a species of marine gastropod mollusc that is one of the biggest sea snails in the Mediterranean, photographed off Vis Island, Croatia, Adriatic Sea
Reef manta ray (Manta alfredi) filter feeding on plankton. Madhivafaru Reef, Raa Atoll, Maldives
Aerial view of West Maui from above Oluwalu, Maui, Hawaii, October 2018
Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina
Common Loons (Gavia immer) one diving underwater, Michigan, USA. June
RF - Oceanic whitetip shark (Carcharhinus longimanus) swims in open waters, close to the surface with sun beams. Egypt. Red Sea. (This image may be licensed either as rights managed or royalty free.)
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Common Frog (Rana temporaria), and frogspawn in a pond, Coldharbour, Surrey, England. March. Highly commended in the Animal Portrait category of the BWPA Competition 2019
Hard coral (Porites lutea) with Christmas-tree worms (Spirobranchus giganteus), New Caledonia, Pacific Ocean
Starling (Sturnus vulgaris) murmuration at sunset, West Pier, Brighton, England, UK
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July
Asian sheepshead wrasse (Semicossyphus reticulatus) mature male in breeding colouration. Niigata, Japan
Killer whale / Orca (Orcinus orca) mature male, swimming underwater. Hamn, Senja, Norway, Atlantic Ocean
Humpback whale calf (Megaptera novaeangliae) male resting beneath his resting mother. Vava u, Tonga, South Pacific
Atlantic walrus (Odobenus rosmarus rosmarus) hauled out on ice floe, Svalbard, Norway June
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Sperm whale (Physeter macrocephalus) female at the ocean surface with her mouth open. This whale has just surfaced from foraging in deep water. Ogasawara or Bonin Islands, Japan
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Alpine bearberry (Arctous alpina) on tundra in autumn and boulderrs at Sydkapp (South Cape), with icebergs in Hall Bredning, Scoresby Sund, Greenland, September
Smalltooth sand tiger shark (Odontaspis ferox), view from below. El Hierro. Canary Islands
Fangtooth moray (Enchelycore anatina) with open mouth, portrait. Tenerife, Canary Islands
Arrow crab (Sternorhynchus lanceolatus). Tenerife, Canary Islands
Short-finned pilot whale (Globicephala macrorhynchus) calf with fetal lines characteristic of newborn specimens, with pod. Tenerife, Canary Islands
Ocean sunfish (Mola mola) and Pilot fish (Naucrates sp). Tenerife, Canary Islands
Napoleon wrasse (Cheilinus undulatus) on a coral reef. Ras Mohammed National Park, Sinai, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
African pompano (Alectis ciliaris) juvenile with long trailing dorsal and anal fins, swimming in mid-water at night. In-camera double exposure
Moon jelly (Aurelia aurita) drifts over a coral reef. East End, Grand Cayman, Cayman Islands, British West Indies. Caribbean Sea
RF - African elephants (Loxodonta africana) in water, trunks touching, Zimanga game reserve, South Africa. (This image may be licensed either as rights managed or royalty free.)
Reef manta ray (Manta alfredi) shoal filter feeding on plankton concentrated by monsoon currents in Raa Atoll, Vandhoo Thila, Maldives. October
Flying fish (Exocoetidae) in Sargasso Sea, Atlantic Ocean