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Spotted tree frog (Litoria spenceri) male, from Still Creek, north-eastern Victoria, Australia. December, Controlled conditions. Critically endangered species
Mountain gorilla (Gorilla beringei beringei) male silverback from Sabyinyo group, Volcanoes National Park, Rwanda, July
Walia ibex (Capra walie) male with Giant lobelia (Lobelia rhynchopetalum) at around 4, 400 meters near Bwahit Pass, Simien Mountains National Park, Amhara, Ethiopia, September
Black Grouse (Lyrurus tetrix) male displaying with bokeh effect in the foreground, Liminka, Finland, March
Portrait of a wild male Bobcat (Lynx rufus), Texas, USA. September
Galapagos penguin (Spheniscus mendiculus) and Marine iguana (Amblyrhynchus cristatus) male, Isla Tortuga, Isabela Island, Galapagos
Red-winged Blackbird (Agelaius phoenicus) male singing from snow-covered red maple (Acer rubrum) in flower, Ithaca, New York, USA. April 2020
Wild Turkey (Meleagris gallopavo) male displaying in early spring, in light snow, New York, USA
Lion-tailed macaque (Macaca silenus), dominant male, Anaimalai Mountain Range (Nilgiri hills), Tamil Nadu, India
Great anglehead lizard (Gonocephalus grandis) male, Tioman Island, Malaysia
Impala (Aepyceros melampus) two males fighting, Mana Pools National Park, Zimbabwe
Lion (Panthera leo) young male, Mana Pools National Park, Zimbabwe
Sand lizard (Lacerta agilis) male on disused railway, wide angle view, the Netherlands
Eurasian bullfinch (Pyrrhula pyrrhula) male perched on twig, Denmark, February
Golden-shouldered parrot (Psephotus chrysopterygius) male landing at nest cavity in termite mound Cape York Peninsula, Queensland, Australia
Adult male lion (Panthera leo) standing on the banks of the Luangwa River, South Lunangwa NP. Zambia
Komodo dragon (Varanus komodoensis) male running. Rinca Island, Komodo National Park, Indonesia. Endangered
Domestic turkey (Meleagris gallopavo) male, close up of iridescence on feathers, France
RF - Red deer (Cervus elaphus) Richmond Park, London, England, UK. (This image may be licensed either as rights managed or royalty free.)
Red deer (Cervus elaphus) silhoutted at dusk, Richmond Park, London, England, UK. October
Dugong (Dugong dugon) male, swims over seagrass meadow (Halophila stipulacea) accompanied by a young Golden trevally (Gnathanodon speciosus). Marsa Nabaa, Marsa Alam, Egypt. Red Sea
Red deer (Cervus elaphus) stag in snowy pine forest. Cairngorms National Park, Highlands, Scotland, UK, March. Highly commended in the Wild Woods Category of the BWPA Awards 2019
Roe deer (Capreolus capreolus) buck in woodland. Peerdsbos, Brasschaat, Belgium. August
Moose (Alces alces) bull, portrait. Grand Teton National Park, Wyoming, USA. September
Moose (Alces alces) bull swimming across river. Grand Teton National Park, Wyoming, USA. September
RF - American bison (Bison bison) male standing near Punch Bowl thermal pool, green algae in foreground. Yellowstone National Park, USA
Puma (Puma concolor puma), young male crossing road in front of car. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December 2018
RF - Skull of a male Himalayan ibex (Capra sibirica) (killed by a snow leopard) lying on steep rocky slope. Himalayas, Ladakh, northern India
RF - Male Violet Sabrewing hummingbirds (Campylopterus hemileucurus) hovering in flight sequence. Montane forest, Bosque de Paz, Caribbean slope, Costa Rica
Growling riflebird (Ptiloris intercedens) male, Adelbert Mountain Range, Madang Province, Papua New Guinea. Endemic
Magnificent Bird of Paradise (Diphyllodes magnificus) male, Lower montane highlands near Mount Hagen, Enga Province, Papua New Guinea. Endemic
Young male Leopard (Panthera pardus) playing with kill after successfully hunting a young Impala at night. South Luangwa National Park, Zambia
Southern humpback whale (Megaptera novaeangliae australis), fluke of male as he dives. Vava'u, Tonga, South Pacific
Playful juvenile Australian sea lion (Neophoca cinerea) greeting me in shallow water at Carnac Island in Western Australia
Desert bighorn sheep (Ovis canadensis nelsoni) rams on steep sandstone wall. Valley of Fire State Park, Nevada, USA. February
RF - Puma (Puma concolor puma), young male walking on hillside near Torres del Paine National Park, Patagonia, Chile. November. (This image may be licensed either as rights managed or royalty free.)
Puma (Puma concolor puma) young male standing with mouth open. Estancia Amarga, near Torres del Paine National Park, Patagonia, Chile. December
Johnston's three-horned chameleon, (Trioceros johnstoni), male on tree branch, Kahuzi-Biega NP, Democratic Republic of Congo, November. Non-ex
Indian leopard (Panthera pardus fusca) on rock with forested hills beyond. Nilgiri Biosphere Reserve, India. Camera trap image. 2019
African wild dog (Lycaon pictus) male, portrait. Beauval Zoo Parc, France. Captive
Giant panda (Ailuropoda melanoleuca) female and juvenile cub aged 2 years, feeding on Bamboo. Yuan Meng was the first Giant panda to be born in France. Zoo Parc de Beauval, France. Captive
Southern double-collared sunbird (Cinnyris chalybeus), male nectaring on Pincushion protea (Leucospermum cordifolium). Cape, South Africa. August
Gold-tailed melitta bee (Melitta haemorrhoidalis), two males sleeping at night in Peach-leaved bellflower (Campanula persicifolia). In garden, Surrey, England, UK. Focus stacked
Malachite sunbird (Nectarinia famosa) male nectaring on King protea (Protea cynaroides). Cape Province, South Africa. August
Fosa (Cryptoprocta ferox) male marking territory onto a mating tree, Kirindy Forest, Madagascar
Red deer, (Cervus elaphus), stag in falling snow on moorland, Scotland, UK. FebruaryRed deer, (Cervus elaphus), stag in falling snow on moorland, Scotland, UK.February
Male and female White-tailed deer (Odocoileus virginianus) with snowman, New York, USA, February
Heath Monitor (Varanus rosenbergi) male enacting a threat display, in mallee / heathHeath Monitor (Varanus rosenbergi) male enacting a threat display, in mallee/heath habitat near Pinnaroo, South Australia
