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Swallowtail butterfly (Papilio machaon) flying to red campion with proboscis extended, Finland, June
Whooper swans (Cygnus cygnus), flying at sunset, Caerlaverock Wildfowl & Wetland Trust, Dumfries & Galloway, Scotland, November 2016
Black-headed gull (Chroicocephalus ridibundus) carrying nesting material, Brenne, France, May
Silver washed fritillary butterfly (Argynnis paphia) female in flight with thistles, Uusimaa, Finland, August
Red-backed shrike (Lanius collurio) adult male taking off, Lower Saxony, Germany, June. Sequence 2 of 3
Southern hawker dragonfly (Aeshna cyanea) flying in habitat, Joutsa, Central Finland, August
Atlantic Puffins (Fratercula arctica) in flight in mist, Skomer Island, Wales, UK, August
Common carder bumblebee (Bombus pascuorum) visiting Lungwort (Pulmonaria officinalis), Monmouthshire, Wales, UK, April
Red mason bee (Osmia bicornis) visiting Geranium sp flower, Monmouthshire, Wales, UK, May
Spotted nutcracker, (Nucifraga caryocatactes) taking off from snow covered tree, Finland, January
Leaf-cutting bee (Megachile species) carrying leaf section. Surrey, England, UK, July
Red-crowned Cranes (Grus japonicus) flying at twilight, Hokkaido, Japan, February
Stellers sea eagle (Haliaeetus pelagicus) in flight with mountains behind, Hokkaido Japan February
Whooper swan (Cygnus cygnus) pair in flight in snowy landscape, Finland, November
Emergence of Mexican free-tailed bats (Tadarida brasiliensis) from Bracken Cave, Texas. Bracken Cave is the summertime home of over 15 million bats
European beewolf (Philanthus triangulum) flying, carrying paralysed bee to nest, Budapest, Hungary
Black headed gull (Chroicocephalus ridibundus) taking off, detail of legs and water droplets, Pusztaszer, Hungary, April
White tailed eagle (Haliaeetus albicilla) in flight, tail feathers caught in morning light, Hokkaido Japan February
Black soldier fly (Hermetia illucens) flying from flower, Tuscaloosa County, Alabama, USA Controlled conditions. September
Sandhill crane (Grus canadensis) landing at sunset. North America. Highly honoured in the Art in Nature category of the Natures Best Photography Windland Smith Rice International Awards Competition
Chinese Sparrowhawk (Accipiter soloensis) flying Guangshui, Hubei province, China, July
Raven (Corvus corax) mobbing Gyrfalcon (Falco rusticolus), Hornoya Island, Varanger Peninsula, Norway, March
Mute swan (Cygnus olor) in flight, Nemunas River Delta, Lithuania, May
Common roller (Coriacias garrulus) in flight with nesting colony behind, Danube delta rewilding area, Romania, June
RF- Eastern white pelicans (Pelecanus onocrotalus) taking off from water, Danube delta rewilding area, Romania. May
White Spoonbill (Platalea leucorodia) in flight, Danube delta rewilding area, Romania May
White tailed sea eagle (Haliaetus albicilla) in flight, Flatanger, Norway, June 2008 DUPE OF 1213183; TO DELETE
RF- Great grey owl (Strix nebulosa) in flight. Kuhmo, Finland
Griffon vulture (Gyps fulvus) landing amongst other vultures on cliff above sea, Liendo Valley within the Montana Oriental Costera, Cantabria, Spain
Beautiful demoiselle damselfly (Calopteryx virgo) female flying, Peerdsbos, Brasschaat, Belgium. June
Griffon vulture (gyps fulvus in flight) Andalusia, Spain, March
Violet sabrewing (Campylopterus hemileucurus) Costa Rica
Puffin (Fratercula arctica) with fish chased by Lesser black-backed gull (Larus fuscus) Isle of May, Firth of Forth, Fife, Scotland, UK, June
Helicopter from Russian icebreaker Kapitan Khlebnikov flies past iceberg in the Weddell Sea near Snow Hill island emperor penguin colony, Antarctica
Demoiselle cranes (Anthropoides virgo) flying at sunrise during migration. Khichan, Western Rajasthan, India. December
Grey-headed lapwing (Vanellus cinereus) winter plumage, flying i, Yangxian Nature Reserve, Shaanxi, China. September
Crested ibis (Nipponia nippon) flying in evening light, Yangxian Nature Reserve, Shaanxi, China. Endangered species. September
Himalayan griffon vulture (Gyps himalayensis) in flight, Dulan Hills, Tibetan Plateau, Qinghai, China
Common vampire bat (Desmodus rotundus) exiting cave, Costa Rica, March 2014
Bewicks swan (Cygnus columbianus) in flight with rainbow, Gloucestershire, England, UK, February
American barn swallow (Hirundo rustica erythrogaster) in flight catching insects over the Gallatin River, Yellowstone National Park, Montana, August
Domestic Fan-tailed pigeons (Columba livia) in flight against a blue sky England, UK
Garden bumblebee (Bombus hortorum), flying to Foxglove (Digitalis purpurea), flower, Monmouthshire, Wales, UK. June
Upland buzzard (Buteo hemilasius), Tibetan Plateau, Qinghai, China
Rufous-tailed hummingbird (Amazilia tzacatl) feeding from Heliconia flower (Heliconia champneiana) La Selva Field Station, Costa Rica
Yellow warbler (Dendroica petechia) collecting nesting material from Bulrush cattail (Typha sp) Bozeman, Montana. USA, June
Small flock of Dunlin (Calidris alpina) with a Sanderling (Calidris alba) flying in front of crashing waves. Druridge Bay, Northumberland, England, UK, January
Snowy owl (Bubo scandiacus) Female taking off the winter tree, Quebec, Canada
