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Great grey owl (Srix nebulosa) in flight about to land in snow, winter, Kuhmo, Finland, February
Greater flamingo (Phoenicopterus roseus) flock standing in lake as others fly overhead and one bird flaps its wings, Camargue, France, January
Grey-headed flying-fox (Pteropus poliocephalus) female, in flight carrying her young, Myuna Wetlands, Doveton, Victoria, Australia
Light-mantled sooty albatross (Phoebetria palpebrata) in flight at sunset, Southern Ocean between Invercargill and Campbell Island, New Zealand, December
Fork-tailed flycatcher (Tyrannus savana) in flight, Pantanal wetlands, Mato Grosso do Sul, Brazil
Blue and yellow macaw (Ara ararauna) in flight, Pantanal, Mato Grosso do Sul, Brazil
Red and green macaw (Ara chloropterus) in flight, Burraco das Araras, Pantanal wetlands, Mato Grosso do Sul, Brazil
White-vented storm petrel (Oceanites gracilis) feeding at sea surface, in flight. Galapagos Islands, Ecuador
Northern gannets (Morus bassanus) breeding colony nesting on sea cliffs, Hermaness, Shetland, Scotland, UK. July
Two Common terns (Sterna hirundo) in middle of aggressive encounter as one lands near nest with chick in it. Long Island, New York, USA. June
Male Downy woodpecker (Picoides pubescens) taking off to avoid Male Hairy woodpecker (Picoides villosus) that landed on its perch in winter, with orange foliage in background
Heermann's gull (Larus heermanni) breeding colony in flight and restless due to presence of pair of Peregrine falcon (Falco peregrinus)
Brown pelicans (Pelecanus occidentalis) flying with Partida and Angel de la Guarda Islands beyond. Rasa Island Special Biosphere Reserve, Sea of Cortez, Mexico. April
The Red Arrows display team flying in formation over the sea, Rhosneigr, Anglesey, Wales, UK. August, 2022
The Royal Air Force Red Arrows display team in formation with red, white and blue contrails, over Rhosneigr, Anglesey, Wales, UK. August, 2022
Common brimstone butterfly (Gonepteryx rhamni) flying towards Purple betony (Betonica officinalis) flowers, Upper Bavaria, Germany. July
Andean condor (Vultur gryphus) soaring above the Three Towers granite rock formation shrouded in cloud, Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Puma (Puma concolor) cub, aged nine months, chasing scavenging White-throated Caracaras (Phalcoboenus albogularis), Torres del Paine National Park / Estancia Laguna Armarga, Patagonia, Chile
Kea (Nestor notabilis) juvenile, taking flight off rock, Arthur's Pass National Park, South Island, New Zealand
Peregrine falcon (Falco peregrinus) female, flying towards nest box with three chicks, aged 18 days, inside, Sagrada Familia Basilica, Catalonia, Spain. May
Galapagos flycatcher (Myiarchus magnirostris) in flight, Santiago Island, Galapagos National Park, Galapagos Islands
Giant hummingbird (Patagona gigas) in flight, portrait, Cotopaxi National Park, Cotopaxi, Ecuador
Split-second prior to Siberian flying squirrel (Pteromys volans orii) landing on tree, adjusting its attitude in mid-flight toward relatively vertical position to reduce speed
Long-tailed tit (Aegithalos caudatus) feeding on icicle formed from sap of Painted maple tree (Acer pictum). Hokkaido, Japan. February
Common starlings (Sturnus vulgaris) large murmuration gathering before landing at winter roost, The Netherlands. Vogelwarte Photo Competition 2022 - Emotion category - Finalist
Mexican violetear hummingbird (Colibri thalassinus) feeding on pink wildflower. Milpa Alta forest, outskirts of Mexico City, Mexico. September
Brent geese (Branta bernicla) small flock in flight over coastal feeding grounds, Lindisfarne National Nature Reserve, Northumberland, UK. February
Greater one-horned rhinoceros (Rhinoceros unicornis) standing in shallow water watching a White-throated kingfisher (Halcyon smyrnensis) fly by, Bardia National Park, Terai, Nepal
Anna's hummingbird (Calypte anna), male flying. Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. November
Common Swift (Apus apus) skimming water surface having just taken a drink. Norwich, UK. July
Red grouse (Lagopus lagopus) in flight over moorland, Yorkshire, UK. September
Black guillemot (Cepphus grylle) holding fish in beak, flying back to nest, Portpatrick, Dumfries and Galloway, Scotland. June
Close up of Galapagos hawk (Buteo galapagoensis) in immature plumage, flying, Alcedo Volcano, Isabela Island, Galapagos Islands, Ecuador
Snowy owl (Bubo scandiaca) female, in flight, Canada. January
Snowy owl (Bubo scandiaca) female, in flight in winter, Canada. January
Flock of Snow geese (Anser caerulescens) leaving their roost at sunrise, Bosque del Apache Wildlife Refuge, New Mexico, USA. January
Ptarmigan (Lagopus mutus) winter plumage, in flight, Utsjoki, Finland. February
Goldcrest (Regulus regulus) in flight, portrait, Parainen Uto, Finland. September
Two Canada geese (Branta canadensis) in flight with autumn foliage behind, Maryland, USA. October
Swallowtail butterfly (Papilio machaon) in flight, Finland, June
Welcome swallow (Hirundo neoxena) in flight over pond. Christchurch, New Zealand
Juvenile Kea (Nestor notabilis) in flight. Arthur's Pass National Park, South Island, New Zealand. Endangered Species
Common Darter (Sympetrum striolatum) in flight, Brasschaat, Belgium
Common tern (Sterna hirundo) adult in breeding plumage flying, carrying fish for young, Long Island, New York, USA, June
Common crane (Grus grus) flying at sunrise, Laguna de Gallocanta, Aragon, Spain
Nankeen kestrel (Falco cenchroides) hovering and looking down, with cresent moon in background, Victoria, Australia
Willow grouse (Lagopus lagopus) winter plumage, in flight, Utsjoki, Finland. February
Redstart (Phoenicurus phoenicurus) juvenile male, in flight, chasing an insect, Parainen Uto, Finland. September
