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Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at dawn. Kasanka National Park, Zambia
Aggregation / flocks of Black Vultures (Coragyps atratus) circling on morning thermals forming over Iguasu Falls, on the Iguasu River, Brazil / Argentina border
Puffin (Fratercula arctica) landing in snow, Norway, March
White capped albatross (Thalassarche steadi) in flight at sea. Auckland Islands (Subantarctic), New Zealand, February
Black-browed albatross (Thalassarche melanophris) in flight over sea at sunrise. South Atlantic Ocean between The Falklands and South Georgia. November
Small tortoiseshell (Aglais urticae) butterfly flying amongst Scabious (Knautia arvensis) in hay meadow. Wensum Valley, Norfolk, England, UK. July
Snow partridge (Lerwa lerwa) flock flying past snowy rockface, in evening light. Jiudingshan Nature Reserve, Mao Country, Sichuan Province, China. November
Eurasian griffon vulture (Gyps fulvus) pre-Pyrenees, Catalonia, Spain. Commended in the Creative Imagery Category of the Bird Photographer of the Year Awards 2019 competition
Bar headed geese (Anser indicus), four in flight, one calling. Coming in to land at breeding site, Qinghai Lake, Qinghai Province, China
Tawny eagle (Aquila rapax) in flight, Zimanga Private Nature Reserve, KwaZulu Natal, South Africa
Tricoloured heron (Egretta tricolor) fishing by flying low over water, amongst Water lilies. Everglades National Park, Florida, USA. March
Glaucous Gull (Larus hyperboreus) taking off from ice, Svalbard Norway, July
Greylag geese (Anser anser) flock in flight and single white plumage bird, Norfolk, UK
Galapagos hawks (Buteo galapagoensis) in flight, Galapagos, Ecuador. Vulnerable species
Magnificent frigatebirds (Fregata magnificens) in low flight ovehead. Espanola Island, Galapagos, Ecuador, December
Hummingbird hawkmoth (Macroglossum stellatarum) Wiltshire, UK, September
Gannet (Morus bassanus) adults and a youngster silhouetted against an orange red sky as another adult prepares to land. Shetland Islands, Scotland, UK. September
Flock of Golden plover (Pluvialis apricaria) in flight, Oare Marshes, Kent, England, UK, September
Flock of Linnets (Carduelis cannabina) in flight over grazing marsh, Elmley Nature Reserve, Kent, England, UK, September
Silhouette of waders flying to roost at sunset, Snettisham RSPB reserve, The Wash, Norfolk, UK, September
Avocets (Recurvirostra avosetta) flock of adults in flight at Snettisham RSPB Reserve. Norfolk, UK, September (non-ex)
Chough (Pyrrhocorax pyrrhocorax) in flight along cliffs, digital composite, South Stack, Anglesey, UK, September
Great bittern (Botaurus stellaris) in flight over water. London, UK. December
Green veined white butterfly (Pieris napi) in flight, Rookery Wood, Sussex, England, UK, August
Territorial Osprey (Pandion haliaetus) chasing a migrant female Marsh Harrier (Circus aeruginosus) from its breeding territory, Manton Bay, Rutland Water, Rutand, UK, August
Gannet (Morus bassanus) in flight is silhouetted against the setting sun. Shetland Islands, Scotland, UK, August
Balearic shearwater (Puffinus mauretanicus) in flight over Irish sea off Pembrokeshire coast, Wales, UK, August 2010, endangered species
Migrant Hawker dragonfly (Aeshna mixta latrielle) male in flight, Wiltshire, England. August
Curlews (Numenius arquata) flying into roost with Hinkley point, Nuclear power station in background, Bridgewater Bay NNR, Somerset, England, August 2009
Northern gannet (Morus bassanus) colony in flight over Bass Rock at sunrise, Firth of Forth, Scotland, August 2009
Small Tortoise-shell butterfly (Aglais urtica) in flight, , with flower, Bavaria, Germany
Pinyon Jay (Gymnorhinus cyanocephalus), gathering Pinyon Pine seeds in autumn. Mono Lake Basin, California, USA, October
Black Skimmer (Rynchops nigra), skimming with its open beak just below the water sBlack Skimmer (Rynchops nigra), skimming with its open beak just below the waters surface in search of prey, Bolsa Chica Ecological Reserve, California
Mustached Bat (Pteronotus sp. ) flying at dusk, Calakmul Biosphere Reserve, Yucatan PeninsulaMustached Bat (Pteronotus sp.) flying at dusk, Calakmul Biosphere Reserve, Yucatan Peninsula, Mexico, August
Himalayan Griffon (Gyps himalayensis) in flight in Spiti valley, Cold Desert Biosphere Reserve, Himalaya mountains, Himachal Pradesh, India, February
Wide angle view of Demoiselle cranes (Anthropoides virgo) landing in chugga ghar (bird feeding enclosure), Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) flock taking flight during winter migration. Khichan, Western Rajasthan, India. February 2015
Mexican free-tailed bats, (Tadarida brasiliensis), leaving Bracken Cave, Texas. Bracken Cave is the summertime home of over 15 milions bats
Great White Egret (Ardea alba) flying above lake blurred motion photograph, Kiskunsagi National Park, Hungary
Juvenile White tailed eagle (Haliaeetus albicilla)in flight, with Hooded crow (Corvus cornix) mobbing it, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Two adult African jacanas (Actophilornis africanus) flying over reedbeds, Selinda Spillway, northern Botswana
White-throated Mountain Gem (Lampornis castaneoventris), Green-crowned Brilliant (Heliodoxa jacula) group of three with one in flight, Savegre, Costa Rica
Downy emerald dragonfly (Cordulia aenea) in flight, Joutsa, Finland, July
Swallowtail butterfly (Papilio machaon) flying to red campion with proboscis extended, Finland, June
Whooper swans (Cygnus cygnus), flying at sunset, Caerlaverock Wildfowl & Wetland Trust, Dumfries & Galloway, Scotland, November 2016
Black-headed gull (Chroicocephalus ridibundus) carrying nesting material, Brenne, France, May
Silver washed fritillary butterfly (Argynnis paphia) female in flight with thistles, Uusimaa, Finland, August
Red-backed shrike (Lanius collurio) adult male taking off, Lower Saxony, Germany, June. Sequence 2 of 3
