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Moths (Lepidoptera) with flight trails, attracted to light during insect trapping and identification session. Long-term monitoring in De Kaaistoep Nature Reserve, Tilburg, The Netherlands. June
Blue jay (Cyanocitta cristata) in flight carrying an acorn that it will cache for winter food supply, New York, USA, October
Osprey (Pandion haliaetus) fishing. Acadia National Park, Maine, USA. May
Great tit (Parus major) male in flight, Parus major, winter, Bavaria, Germany, February
Great Grey Owl (Strix nebulosa) flying directly overhead, Kuhmo Finland February
Monarch butterflies (Danaus plexippus) flying during a warm morning. When the overwintering period draws to an end large numbers become active. Michoacan, Mexico, February
Honey bee (Apis mellifera) approaching hive entrance, with trails of departing bees visible. Urban garden, Bristol, UK, May
Honey bee (Apis mellifera) flying close to hive entrance, with flight trails visible. Urban garden, Bristol, UK, May
RF - Red kite (Milvus milvus) in flight. Calera y Chozas, Castile-La Mancha, Spain. December. (This image may be licensed either as rights managed or royalty free.)
Peacock butterfly (Aglais io) flying over flowers. Finland. August
Siberian jay (Perisoreus infaustus) flying in coniferous forest. Kuusamo, Northern Ostrobothnia, Finland. February
Apollo (Parnassius apollo) butterfly female in flight over field, sphragis present following copulation. Pargas, Aboland, Finland. July 2019
Schneiders leaf-nosed bats (Hipposideros sperosis) in half submerged old temple ruins. Hampi World UNESCO Heritage Site, Karnataka, India
Naked-rumped tomb bats (Taphazhopus theobaldi), flying out of Garbhgriha stone carved door of 6th century AD old Shiv temple. Badami, Karanataka, India
Dense flock of Common teal (Anas crecca) with a few Wigeon (Anas penelope) and Northern shoveler (Anas clypeata) in flight over others resting on partly frozen flooded marshes
Ruppells griffon vulture (Gyps rueppellii) flying over Jinbar waterfall, Simien Mountains National Park, Amhara, Ethiopia, September. Critically endangered species
Waved albatross (Phoebastria irrorata) in flight, Punta Suarez, Espanola Island, Galapagos
Brown pelican (Pelecanus occidentalis) landing reflected in tide pool, Puerto Egas, Santiago Island, Galapagos
Brown pelicans (Pelicanus occidentalis) flock in flight over sea, Galapagos
Swallow-tailed gull (Creagrus furcatus) in flight, Punta Cevallos, Espanola Island, Galapagos
Red-footed booby (Sula sula) landing among salt bushes, looking for nesting materials, with Frigate birds behind, Genovesa Island, Galapagos
Galapagos hawk (Buteo galapagoensis) in flight, Alcedo Volcano, Isabela Island, Galapagos
Nazca booby (Sula granti) in flight at dusk. Punta Suarez, Espanola Island, Galapagos
Blue-footed booby (Sula nebouxii) on sea, tossing fish. Turtle Cove, Santa Cruz Island, Galapagos
Galapagos carpenter bee (Xilocopa darwini) visiting native passionflower (Passiflora foetida) in photographer Tui De Roys garden, Santa Cruz Island, Galapagos Islands
Orchid bee (Euglossa sp. ) visits an orchid in cloud forest, Choco regionOrchid bee (Euglossa sp.) visits an orchid in cloud forest, Choco region, Northwestern Ecuador
Velvet-purple coronet hummingbird (Boissonneaua jardini) visits a flower in cloud forest, Choco region, Northwestern Ecuador
Southern hawker (Aeshna cyanea) dragonfly in flight, Broxwater, Cornwall, UK. August
RF - Bar-tailed godwit (Limosa lapponica) in flight over the Wash, Norfolk, UK. November. (This image may be licensed either as rights managed or royalty free.)
RF - Painted lady butterfly (Cynthia cardui) feeding on Fleabane (This image may be licensed either as rights managed or royalty free.)
Laughing Gull (Larus atricilla) flock in flight, Sisal, Yucatan Peninsula, Mexico, January
Common cranes (Grus grus) leaving the roost at dawn, Hula Valley, Northern Israel, January
Atlantic puffin (Fratercula arctica) in breeding plumage flapping wings on sea cliff top in seabird colony, Scotland, UK, May. Digital composite
Great skua (Stercorarius skua) in flight and spectacular coastline with sea stacks and cliffs, home to breeding sea birds at Hermaness, Unst, Shetland Islands, Scotland. May 2018. Digital composite
Red-throated Diver (Gavia stellata) taking off with fish prey, Iceland, June
Hawk Owl (Surnia ulula) hunting, Kuhmo, Finland, March
White-tailed Eagle (Haliaetus albicilla) Norway, November
Honey garlic (Nectaroscordium siculum) with Bumblebee in flight. Naturalised in the Avon Gorge, Bristol, UK, June
Monarch butterfly (Danaus plexippus) wintering in Oyamel pine forests (Abies religiosa) Monarch Butterfly Biosphere Reserve / Reserva de Biosfera de la Mariposa Monarca UNESCO World Heritage Site
Northern fulmar (Fulmarus glacialis) in flight with waves, Arctic Sea. Commended in the Biophoto Competition 2019
African elephant (Loxodonta africana) two silhouetted at sunset with goose flying overhead, Mkuze, South Africa. Highly commended in the African Wildlife category of the Nature's Best Photography
Crab-plover (Dromas ardeola) flying with crab in beak. Oman, June
Crab-plover (Dromas ardeola) three flying, two with crabs in beaks. Oman, June
Red-and-green macaws (Ara chloropterus) pair in flight over forest canopy. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
RF - Andean condor (Vultur gryphus) flying over Torres del Paine Massif. Torres del Paine National Park, Chilean Patagonia, Chile. March
Group of Red-and-green macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Straw-coloured fruit bats (Eidolon helvum) returning to daytime roost at dawn. Kasanka National Park, Zambia
