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Nutcracker (Nucifraga caryocatactes) in flight with wings spread, Sipoo, Finland. January
Wood sandpiper (Tringa glareola) in flight, Parainen Uto, Finland. August
Hen harrier (Circus cyaneus) juvenile male, in flight, Parainen Uto, Finland. September
Great egret (Ardea alba) flying with stick for its nest in beak. St. Augustine, Florida, USA
Great blue heron (Ardea herodias) flying over foggy river at sunrise. Myakka River State Park, Florida, USA. Sequende 2 of 2
Red kite (Milvus milvus) landing on grass, Marlborough Downs, Wiltshire, UK. January
Male Giant kingfisher (Megaceryle maxima) diving, Allahein River, The Gambia
Flock of White-faced whistling duck (Actophilornis africanus) in flight, Allahein River, The Gambia
Mute swans (Cygnus olor) in flight against a cloudy sky, Caerlaverock Wetland Centre, Dumfries & Galloway, Scotland, UK. November
Grey-headed flying-fox (Pteropus poliocephalus) in flight, with tongue out after licking water from its body. Having just completed a high-speed belly dip into a river on a hot day to wet its belly
Arctic tern (Sterna paradisaea) in flight over nesting area in meadows near coast, Reykjanes Peninsula, Iceland. July
Little owl (Athene noctua) taking flight, NorthYorkshire, UK. June, 2021
Eurasian hoopoe (Upupa epops) in flight with a grub in its beak, Pusztaszer reserve, Hungary. May
European roller (Coracias garrulus) in flight, Pusztaszer reserve, Hungary. May
European turtle dove (Streptopelia turtur) in flight, Pusztaszer reserve, Hungary. May
Broad-bordered bee hawk-moth (Hemaris fuciformis) nectaring from Wood cranesbill (Geranium sylvaticum), Finland. June
Adult White-tailed eagle (Haliaeetus albicilla) taking off from its perch, Finland. July
Wood warbler (Phylloscopus sibilatrix) taking off flowering willow, Finland. May
Flock of Sanderlings (Calidris Alba) in flight with wind turbines in background, Atlantic Coast, Vendee, France. December
Black tern (Chlidonias niger) hunting over water in prairie wetland for emerging midges (Chironomidae) Medicine Lake National Wildlife Refuge, Montana, USA. June
Palm-nut vulture (Gypohierax angolensis) in flight, Allahein river, The Gambia
Small flock of migrating Black-tailed godwits (Limosa limosa) in flight against a bright blue sky, River Coquet Estuary, Northumberland, UK. September
Jackdaw (Corvus monedula) pair dive bombing and harassing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel
Jackdaw (Corvus monedula) swooping down calling with claws raised to threaten a Grey squirrel (Sciurus carolinensis) in the entrance to a nest box the bird wants to nest in which is already occupied
Jackdaw (Corvus monedula) swooping in very close to threaten a Grey squirrel (Sciurus carolinensis) as it stands in the entrance to a nest box the bird wants to nest in which is already occupied by
Marbled white butterfly (Melanargia galathea) sequence showing landing and taking off from a Greater knapweed (Centaurea scabiosa) flower. Yorkshire, UK. Digital composite
A female kingfisher (Alcedo atthis) fishing / diving into an ice hole in winter. Leeds, Yorkshire, UK. January
Ruby-throated hummingbird (Archilochus colubris) female nectaring on Maximilian sunflower (Helianthus maximilianii). Hill Country, Texas, USA
Kingfisher (Alcedo atthis) male, after diving, taking off from water, Lorraine, France, July
Tawny eagle (Aquila rapax) in flight, coming into land on carcass. Ndutu, Ngorongoro Conservation Area / Serengeti Tanzania
Male Violet Sabrewing (Campylopterus hemileucurus) hovering / in flight sequence. Montane forest, Bosque de Paz, Caribbean slope, Costa Rica, Central America
Ventral view of Swallowtail butterfly (Papilio machaon) in flight. The Netherlands. July
RF - Two Swallowtail butterflies (Papilio machaon) flying to feed on thistle (Cirsium sp. ) bush in flowerRF - Two Swallowtail butterflies (Papilio machaon) flying to feed on thistle (Cirsium sp.) bush in flower. The Netherlands. July
Painted lady butterfly (Vanessa cardui) taking flight, Norfolk, England, UK. June
Galah (Eolophus roseicapillus) flock of birds flying over grassland, Diamantina River, Queensland, Australia
Male Great frigatebird (Fregata minor) in breeding condition with semi-inflated throat pouch, Christmas Island, Australia
Orange leaf-nosed bat (Rhinonicteris aurantia) flying out of abandoned mine in late evening, Pine Creek, Northern Territory, Australia
Hills sheathtail bat (Taphozous hilli) flying out of an abandoned mine in late evening, Tennant Creek, Northern Territory, Australia
Eastern horseshoe bat (Rhinolophus megaphyllus) flying out from an abandoned mine in late evening. Its orange colouring is due to bleaching from the intense ammonia atmosphere in the mine
Cranefly (Tipula paludosa) flight trail over meadow in old quarry on autumn night in Somerset, England. September
Caribbean flamingos (Phoenicopterus ruber) flying, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Common tern (Sterna hirundo) catching a mayfly, Hjalstaviken, Uppland, Sweden. Nordic Nature Photo Competition 2021 Highly Commended - Nordic Bird Photo of the Year category
Black stork (Ciconia nigra) in flight during spring migration across the Sahara desert, Aydar, Southern Morocco, Africa
Great grey owl (Strix nebulosa) preparing to land, in tree, Yellowstone National Park, Wyoming, USA
A female kingfisher (Alcedo atthis) fishing / diving into an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence 1 of 4
A female kingfisher (Alcedo atthis) fishing, flying out of an ice hole in winter. Leeds, Yorkshire, UK. January
Firecrest (Regulis ignicapilla) in flight, Parainen, Uto, Finland. October
Golden-tailed sapphire (Chrysuronia oenone) in flight. Pilcopata, Cuzco, Peru. May