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Puffins (Fratercula arctica) pair take off together to go fishing. Farne Islands, Northumberland, UK, May. Non-ex
Great Tit (Parus major) in flight. Bavaria, Germany, December
A pair of Toco Toucans (Ramphastos toco) flying across the Piquiri River at dawn, northern Pantanal, Mato Grosso, Brazil, September. (Digitally Modified)
Snowy Owl (Bubo scandiaca) in flight over snow covered landscape, Ottawa, Ontario, Canada, December
Violet Sabrewing Hummingbird (Campylopterus hemileucurus) male hovering at feeder, Bosque de Paz, Costa Rica, Central America
Arctic tern (Sterna Paradisaea) hovering in flight over an iceberg. Peterman IslandArctic tern (Sterna Paradisaea) hovering in flight over an iceberg.Peterman Island, Antarctic Peninsula, Antarctica. February
Flock of Kelp / Southern black backed Gulls (Larus dominicanus) flying over iceberg. Graham Passage, Antarctic Peninsula, Antarctica. February. (digitally altered image)
Green woodpecker {Picus viridis} male in flight, Lorraine, France
Flock of Sanderling (Calidris alba) flying over a rough sea, Berwickshire Coast, Scotland, UK, March
Black browed albatross flying over iceberg {Thalassarche melanophrys} South Georgia
Bullers albatross (Thalassarche bulleri) flying over sea, Chatham Islands, off southern New Zealand
Common starling (Sturnus vulgaris) flock flying towards their night time roost, Scottish Borders, Scotland, UK
Sword-billed Hummingbird (Ensifera ensifera) female feeding from Datura flower, Papallacta, Ecuador, Andes, South America, January
Ruby-topaz Hummingbird (Chrysolampis mosquitus) male flying demonstrating how brilliant colours only show when the bird is viewed from the front, Digital composite, Trinidad
Goshawk (Accipiter gentilis) in flight over woodland clearing, Cairngorms National Park, Scotland, UK
Ferruginous hawk (Buteo regalis) landing. Captive
Feral pigeon (Rock Dove) flying, UK
Fulmar (Fulmarus glacialis) rear view of bird hanging in air over steep cliffs, Shetland Islands, Scotland, UK. HIGHLY COMMENDED - Animals in their environment - Wildlife Photographer of the Year
Light-mantled sooty albatross (Phoebetria palpebrata) in flight at sunset, Southern Ocean between Invercargill and Campbell Island, New Zealand, December
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
White-tailed eagle (Haliaeetus albicilla) in flight over mountain landscape at dusk, Iceland, January
Whooper swan (Cygnus cygnus) in flight over frozen lake at dusk, Lancashire, UK December
Rock Martin (ptyonoprogne fuligula)in flight, Israel
Crested terns (Thalasseus bergii) flying over a coastal rock shelf in the early morning with waves breaking in the background. Long exposure. Skenes Creek, Victoria, Australia. July 2016
Great grey owl (Strix nebulosa) taking off from snow, Kuhmo, Finland, February
Japanese / Red-crowned crane (Grus japonicus) three in flight, Hokkaido Japan February
A flock of Starlings (Sturnus vulgaris) flying in front of the Blackpool Tower, England, UK, September 2010
Wandering Albatross (Diomedea exulans) flying over the rugged coast near Stromness, South Georgia, South Atlantic. (digitally stitched image) February
Oystercatcher (Haematopus ostralegus) in flight over wildflowers, Great Saltee, Saltee Islands, County Wexford, Republic of Ireland. May
A female kingfisher (Alcedo atthis) fishing through an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Scarlet Macaw (Ara macao) couple in flight and breaking to land Corcovado National Park, Osa peninsula, Costa Rica
Grey heron (Ardea cinerea cinerea) in flight at last light, Blackpool, UK, February
Sequence of a Red-and-green macaw or green-winged macaw (Ara chloropterus) coming into land at its nest hole on a cliff. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
RF - Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Common buzzard (Buteo buteo) in flight over snow, Pusztaszer, Hungary, January
Greater flamingos (Phoenicopterus ruber) in flight at dawn, Camargue, France. Digitally added flamingo
Light-mantled Sooty Albatross (Phoebetria palpebrata) flying over Fortuna Glacier with rainbow, Fortuna Bay, South Georgia. January 2010. (digitally stitched image)
Black-browed Albatross (Thalassarche melanophrys) flying off the Bay of Isles, South Atlantic. January.(digitally stitched image)
Egyptian goose (Alopochen aegyptiaca) in flight, London, UK, February
Caribbean flamingo (Phoenicopterus ruber) landing, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Aerial view of Caribbean flamingos (Phoenicopterus ruber) flying over the coastal lagoon, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
Bald eagle (Haliaeetus leucocephalus) flying over water in falling snow. Kachemak Bay, Alaska, USA. February
Large flock of Guillemots (Uria aalge) taking off from a cliff in falling snow, Vardo, Norway, March
Ventral view of Swallowtail butterfly (Papilio machaon), male and female, in flight. The Netherlands. July
Caribbean flamingo (Phoenicopterus ruber) flying, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January
Kittiwakes (Rissa tridactyla) flying over surf, Vardo, Norway March
Great grey owl (Srix nebulosa) in flight about to land in snow, winter, Kuhmo, Finland, February