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Bearded vulture (Gypaetus barbatus) flying, Ordesa National Park, Pyrenees, Spain
Common swift (Apus apus) flying low to water while hunting, Norfolk, England, UK, July. Winner of Animal Behaviour Category of the British Wildlife Photography Awards 2019. Photographer
White-tailed sea eagle (Haliaeetus albicilla) flying over water with male Common eider prey (Somateria mollissima), Uto, Parainen, Finland, April
Male Black grouse (Tetrao / Lyrurus tetrix) in flight. Kuusamo, Finland, April
Buzzard (Buteo buteo) landing on branch, winter, Lorraine, France, February
Kingfisher (Alcedo atthis) male, after diving, taking off from water with fish, a Common Roach (Rutilus rutilus) Lorraine, France, July
White-tailed eagle (Haliaeetus albicilla) flying over water with fish in talons. Norway. August
Great grey owl (Strix nebulosa) in flight in woodland during snowfall. Kuusamo. Finland. February
Griffon vulture (Gyps fulvus) soaring over Los Mallos de Riglos, Huesca province, Aragon, Spain
African fish eagle (Haliaeetus vocifer) swoops to catch a freshly caught fish, dropped by a Saddle-billed stork (Ephippiorhynchus senegalensis) after being pressurised to do so by the eagle
Taiwan blue magpie, (Urocissa caerulea) flying, in Taipeh, Taiwan, endemic species
Peregrine falcon (Falco peregrinus) in flight with open beak. The Netherlands. May
California condor (Gymnogyps californianus) landing, forest in background. Bird with tags on wings. Near San Pedro Martir National Park, Northern Baja California, Mexico. 2017
RF - Red-and-green macaws (Ara chloropterus) two with one taking off, Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
European bee eaters (Merops apiaster) one taking off another perched, Kiskunsagi National Park, Pusztaszer, Hungary. May
Red-and-green macaw (Ara chloropterus) in flight, Pantanal, Brazil. August
Crested tit (Lophophanes cristatus) about to land on Elmleaf blackberry (Rubus ulmifolius) plant, Sierra de Grazalema Natural Park, Southern Spain, January
Common tern (Sterna hirundo) flying low over water. Hjalstaviken, Uppland, Sweden
Great white egret (Egretta / Ardea alba) Pusztaszer protected landscape, Kiskunsagi, Hungary, May
Blue jay (Cyanocitta cristata) flying, head on, Acadia National Park, Maine, USA. May
Bullfinch (Pyrrhula pyrrhula) in flight, Kuusamo, Finland, March
Black-veined white (Aporia crataegi) butterfly pair, in flight and nectaring on Thistle. Jyvaskyla, Central Finland. June
Pin-tailed whydah (Vidua macroura) male in display flight. Allahein River, Gambia
RF - Atlantic Puffins (Fratercula arctica) flying on to the cliff top and gathering around burrows. Isle of Lunga, Treshnish Isles, Isle of Mull, Scotland
Gila woodpecker (Melanerpes uropygialis), emerging from nest in Saguaro cactus, Arizona, USA. July
Marsh harrier (Circus aeruginosus) female hunting among a panicked flock of Common teal (Anas crecca) in flight over marshland, RSPB Greylake Nature Reserve, Somerset, UK, December
Atlantic puffin (Fratercula arctica) taking off, Machias Seal Island, Maine, USA, June
Common tern (Sterna hirundo) close up in flight, Tenerife, Canary Islands
Galapagos hawk (Buteo galapagoensis) flying, carrying an iguana carcass, Punta Espinosa, Fernandina Island, Galapagos
Short-eared owl (Asio flammeus) in flight, Santa Fe Island, Galapagos
Buff-tailed bumblebee (Bombus terrestris) queen about to land at her nest burrow in a garden lawn with loaded pollen sacs to provision grubs that will become future workers for her colony, Wiltshire
Firefly (Lamprohiza splendidula) light trails of males in forest, Bavaria, Germany. July
Golden eagle (Aquila chrysaetus) landing in snow, Kuusamo, Finland, December
Dipper (Cinclus cinclus) taking off from water, Kuusamo, Finland, February
Black Woodpecker male (Dryocopus martius) Finland, January. One branch removed
Great white pelican (Pelecanus onocrotalus) flying over water, Danube Delta, Romania. July
Cuckoo (Cuculus canorus) in flight, Germany, April. May
Red-and-green macaw (Ara chloropterus) flock feeding at wall of clay lick, flying and perched in trees. Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Greater flamingo (Phoenicopterus roseus) legs of bird in flight, Pont Du Gau Park, Camargue, France
Scopolis shearwater (Calonectris diomedea) in flight. Tenerife, Canary Islands
Brunnichs guillemot (Uria lomvia) colony in mist, many birds in flight and nesting on cliffs. Svalbard, Norway, July 2018
Barn Owl (Tyto alba) landing on gravestone, captive, digital composite, UK
Magnificent frigatebird (Fregata magnificens) in flight over sea between Isabela Island and Fernandina Island, Galapagos
California condor (Gymnogyps californianus) in flight, wings radio tagged. California condor recovery program, Sierra de San Pedro Martir National Park, Baja California Peninsula, Mexico. 2011
Purple throated mountain gem hummingbird (Lampornis calolaemus) hovering as it nectars on Porterweed (Stachytarpheta frantzii). Costa Rica
Rhinoceros hornbills (Buceros rhinoceros) pair flying over the canopy with mist hanging over lowland Dipterocarp rainforest in heart of Danum Valley, Sabah, Borneo (digitally modified image)
Short-toed Eagle (circaetus gallicus) flying amongst Stone Pines (Pinus pinea). Coto Donyana, Andalusia, Spain. 2007
