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Garden bumblebee (Bombus hortorum) flying to Hedge woundwort (Stachys sylvatica) Monmouthshire, Wales, UK, July
Common Swift (Apus apus) in flight reflected in water, Norfolk, England, UK. July
Little bittern (Ixobrychus minutus) watching Great warbler (Acrocephalus arundinaceus) with prey, Lake Csaj, Kiskunsagi National Park, Hungary
Two coloured mason bee (Osmia bicolor) bee that nests in old snail shells and then carries bits of twig or vegetation to cover the shell, Buckinghamshire, England, UK, June
Copper-rumped hummingbird (Saucerottia tobaci) feeding, Tobago, West Indies
Flock of Mallard ducks (Anas platyrhynchos) taking off, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. December
Common raven (Corvus corax) two adults in flight, Hornoya, Norway, April
Eurasian griffon vulture (Gyps fulvus) in flight, Cevennes, France, March 2016
European Greenfinch (Chloris chloris) flying at dawn. When backlit the tiny transparent barbs that make up the feather act like a prism revealing vivid colours in the early light
Brown booby (Sula leucogaster), adult diving for fish, Los Roques, Venezuela
Series of Bald eagle (Haliaeetus leucocephalus) in flight, Homer, Alaska, USA, March. Digital composite
Great Frigatebird (Fregata minor) arriving at nest with chick, San Benedicto Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site
Rock ptarmigan (Lagopus muta) in flight, Utsjoki, Finland, April
Blue cheeked bee eater (Merops persicus) two squabbling, Oman, April
Siskin (Carduelis spinus) male in flight preparing to land, Scotland, UK, May
Gannet (Morus bassanus) coming in to land in the colony in evening light, Great Saltee, Saltee Islands, County Wexford, Ireland, June
Pink-footed goose (Anser brachyrhynchus) flock flying in mist, Cromarty Firth, Highlands, Scotland, UK April
Hoopoe (Upupa epops) male in flight, Hungary May
Green lacewing (Abachrysa eureka) in flight, , Tuscaloosa County, Alabama, USA Controlled conditions. May
Harlequin ladybird (Harmonia axyridis) flying, Tuscaloosa County, Alabama, USA Controlled conditions. January
Red wasp (Polistes carolina) in flight Lamar County, Texas, USA Controlled conditions. August
Hawk owl (Surnia ulula) takingn off from powder snow, possibly with prey in talons, Kuusamo Finland February
Killer whale (Orcinus orca) adult male surfacing at dusk surrounded by birds, who watch for herrings (Clupea harengus) left after they have fed, Andenes, Andoya island, North Atlantic Ocean, Norway
Black skimmer (Rynchops niger) foraging along surf line. Tierra Verde, Florida, USA
RF - Red-and-green macaw (Ara chloropterus) flock feeding and flying. Wall of clay lick, Heath River, Tambopata / Bahuaja-Sonene Reserves, Amazonia, Peru / Bolivia border
Hooded crown (Corvus cornix) flying with egg in beak, Danube Delta, Romania. May
Kittiwake (Rissa tridactyla) flying over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Herring gull (Larus argentatus) flying over sea. Great Saltee Island, Saltee Islands, County Wexford, Republic of Ireland. June
Puffin (Fratercula arctica) in flight, Saltee Islands, County Wexford, Republic of Ireland, June
Northern gannet (Morus bassanus), Feeding frenzy, Noss NNR, Shetland, Scotland, UK
Snowy owl (Bubo scandiacus) in flight, Wrangel Island, Far Eastern Russia, October
Wild Konik horses with Starlings (Sturnus vulgaris) Odry delta reserve, Stepnica, Poland, July
Low angle view of Demoiselle crane (Anthropoides virgo) flying and feeding during winter migration. Khichan, Western Rajasthan, India. December
Demoiselle crane (Anthropoides virgo) low angle view of birds flying and landing near the chugga ghar (bird feeding enclosure ) in Khichan Village, durin their annual migration
Wreathed hornbill (Aceros undulatus) taking off, Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China. April
Spoonbill (Platalea leucorodia) carrying nest material, The Netherlands
Mining bee (Andrena sp) flying to Giant harebell (Campanula latifolia), Pentwyn farm SSSI, Gwent Wildlife Trust Reserve, Monmouthshire Wales UK, July
Saker falcon (Falco cherrug milvipes) in flight, Keke Xili, Changtang, Tibetan Plateau, Qinghai, China
Common wasp (Vespula vulgaris) flying to fuchsia flower, Banbridge, County Down, Northern Ireland
RF - Brunnichs guillemot (Uria lomvia) group flying in formation, Svalbard, Norway
Kea (Nestor notabilis) juvenile in flight. Arthurs Pass National Park, South Island, New Zealand. Endangered Species
Westland petrel (Procellaria westlandica) in flight over sea, showing the upperwing. Kaikoura, South Island, New Zealand. April. Vulnerable species
Golden pheasant 1+Chrysolophus pictus+2 male jumping, Yangxian Biosphere Reserve, Shaanxi, China
Snowy owl (Bubo scandiacus) Female landing in the snow, Quebec, Canada
Crested ibis (Nipponia nippon) flying, Yangxian Nature Reserve, Shaanxi, China, September
Oriental Pied hornbill (Anthracoceros albirostris) male landing at nest hole, Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Great hornbill (Buceros bicornis) in flight, blurred motion. Tongbiguan Nature Reserve, Dehong Prefecture, Yunnan Province, China, April
Red-billed blue magpie (Urocissa erythroryncha) flying, Yangxian Biosphere Reserve, Shaanxi, China