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Golden eagle (Aquila chrysaetus) landing, Kuusamo, Finland, January
Siberian Jay (Perisoreus infaustus) alighting on a branch. Hamra, Sweden, September
Two Common ravens (Corvus corax) in flight through snow, Finland, April
Osprey (Pandion haliaetus) flying through a rainbow. Aviemore, Scotland, July
Yellowhammers (Emberiza citrinella) coming in to land of a sheaf of oats in winter. Perthshire, Scotland, February
Purple-bibbed Whitetip (Urosticte benjamini) male feeding from flower, Mindo, Ecuador, Andes, South America, January
Green Woodpecker (Picus viridis) male flying through Beech woodland in spring, digital composite, Surrey, UK
Barn owl {Tyto alba} female landing on fence post. Captive UK
Great egret (Ardea alba) flying across lake, Ranthambhore National Park, India
Atlantic Puffin (Fratercula arctica) in flight, coming into land with beak full of sand eels (Ammodytes tobianus). Fair Isle, Shetland Islands, Scotland, UK, July
Sword billed hummingbird (Ensifera ensifera) in flight, Guango, Napo, Ecuador
Light-mantled sooty albatross (Phoebetria palpebrata) in flight at sunset, Southern Ocean between Invercargill and Campbell Island, New Zealand, December
Caribbean flamingo (Phoenicopterus ruber) flock flying in formation, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
White-tailed eagle (Haliaeetus albicilla) in flight over mountain landscape at dusk, Iceland, January
Whooper swan (Cygnus cygnus) in flight over frozen lake at dusk, Lancashire, UK December
Rock Martin (ptyonoprogne fuligula)in flight, Israel
Crested terns (Thalasseus bergii) flying over a coastal rock shelf in the early morning with waves breaking in the background. Long exposure. Skenes Creek, Victoria, Australia. July 2016
Great grey owl (Strix nebulosa) taking off from snow, Kuhmo, Finland, February
Japanese / Red-crowned crane (Grus japonicus) three in flight, Hokkaido Japan February
A flock of Starlings (Sturnus vulgaris) flying in front of the Blackpool Tower, England, UK, September 2010
Wandering Albatross (Diomedea exulans) flying over the rugged coast near Stromness, South Georgia, South Atlantic. (digitally stitched image) February
Oystercatcher (Haematopus ostralegus) in flight over wildflowers, Great Saltee, Saltee Islands, County Wexford, Republic of Ireland. May
A female kingfisher (Alcedo atthis) fishing through an ice hole in winter. Leeds, Yorkshire, UK. January. Sequence
Male Black grouse (Tetrao / Lyrurus tetrix) displaying at lek. Kuusamo, Finland, April
Scarlet Macaw (Ara macao) couple in flight and breaking to land Corcovado National Park, Osa peninsula, Costa Rica
Grey heron (Ardea cinerea cinerea) in flight at last light, Blackpool, UK, February
Sequence of a Red-and-green macaw or green-winged macaw (Ara chloropterus) coming into land at its nest hole on a cliff. Buraco das Araras, Jardim, Mato Grosso do Sul, Brazil. September
RF - Red-and-Green Macaws or Green-winged Macaws (Ara chloropterus) in flight over forest canopy. Buraco das Araras (Sinkhole of the Macaws), Jardim, Mato Grosso do Sul, Brazil. September
Common buzzard (Buteo buteo) in flight over snow, Pusztaszer, Hungary, January
Greater flamingos (Phoenicopterus ruber) in flight at dawn, Camargue, France. Digitally added flamingo
Light-mantled Sooty Albatross (Phoebetria palpebrata) flying over Fortuna Glacier with rainbow, Fortuna Bay, South Georgia. January 2010. (digitally stitched image)
Black-browed Albatross (Thalassarche melanophrys) flying off the Bay of Isles, South Atlantic. January.(digitally stitched image)
Egyptian goose (Alopochen aegyptiaca) in flight, London, UK, February
Caribbean flamingo (Phoenicopterus ruber) landing, Ria Celestun Biosphere Reserve, Yucatan Peninsula, Mexico, January. Bookplate
Aerial view of Caribbean flamingos (Phoenicopterus ruber) flying over the coastal lagoon, Ria Lagartos Biosphere Reserve, Yucatan Peninsula, Mexico, May. Bookplate
Bald eagle (Haliaeetus leucocephalus) flying over water in falling snow. Kachemak Bay, Alaska, USA. February
Large flock of Guillemots (Uria aalge) taking off from a cliff in falling snow, Vardo, Norway, March
Ventral view of Swallowtail butterfly (Papilio machaon), male and female, in flight. The Netherlands. July
Japanese dwarf flying squirrel (Pteromys volans orii) gliding through forest canopy during light snowfall. Hokkaido, Japan. February
Jackdaw (Corvus monedula) pair chasing a Grey squirrel (Sciurus carolinensis) as it emerges from a nest box the birds want to nest in which is already occupied by the squirrel and its mate
Jackdaw (Corvus monedula) hovering above a nest box it wants to nest in which a Grey squirrel (Sciurus carolinensis) and its mate have occupied, with the squirrel threatening the bird, Wiltshire, UK
Ghost bat (Macroderma gigas) in flight, Pine Creek, Northern Territory, Australia
Sparrowhawk (Accipiter nisus) juvenile flying, Parainen Uto, Finland. September
Goshawk (Accipiter gentilis) flying in snow, Mainua Kajaani, Finland. March
Annas hummingbird (Calypte anna) male dancing in flight and displaying his bright pink throat feathers to a female, Montana, USA. June
Firefly (Lamprohiza splendidula) light trails of males in forest at dusk, Bavaria, Germany. July
Dusky leaf-nosed bat (Hipposideros ater) flying out of daytime cave roost in a sandstone escarpment, Kimberley, Western Australia
Bearded vulture (Gypaetus barbatus) flying, Ordesa National Park, Pyrenees, Spain