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White-rumped Snowfinch (Montifringilla taczanowskii) with chick at nest burrow, Sanjiangyuan National Nature Reserve, Qinghai Hoh Xil UNESCO World Heritage Site, Qinghai-Tibet Plateau
Silver Birch (Betula pendula) in autumn, Craigellachie National Nature Reserve, Speyside, Cairngorms National Park, Scotland, October
Beech (Fagus sylvatica) mature tree in winter, Paxton House Estate, Berwickshire, Scotland, December
Woodcock (Scolopax rusticola) probing for invertebrate prey in marsh in wintry conditions, Berwickshire, Scotland, January
Whooper Swan (Cygnus cygnus) male on moorland loch, Isle of Skye, Hebrides, Scotland, December
Reindeer (Rangifer tarandus) bull reindeer with antlers in velvet, reintroduced Cairngorm Reindeer Herd, Cairngorm National Park, Speyside, Scotland, October
Borage flower bud (Borago officinalis) near Nice, south of France, June
Lance-nosed chameleon (Calumma gallus), Andasibe-Mantadia National Park. Madagascar
Weaver ants (Oecophylla smaragdina) holding leaves together during nest building, Malaysian Borneo
White tent making bat (Ectophylla alba) roosting in tree, La Selva Field Station, Costa Rica
Lesser celandine (Ranunculus ficaria) Clare Glen, Tandragee, County Armagh. Northern Ireland. March
Cartwheeling spider (Carparachne sp. ) in desert, Swakopmund, NamibiaCartwheeling spider (Carparachne sp.) in desert, Swakopmund, Namibia
Weaver ants (Oecophylla smaragdina) portrait, Sabah, Malaysian Borneo
Dandy jumping spider (Portia schultzi) Kwazulu-Natal, South Africa
Stormy sky above the prairie grassland, Montana, USA. June 2011
Common wasp (Vespula vulgaris) flying to fuchsia flower, Banbridge, County Down, Northern Ireland
Amethyst deceiver (Laccaria amethystina) fungus, Peatlands Park Co. Armagh, Northern Ireland
Giant African mantis (Sphodromantis viridis) portrait, captive, occurs in West Africa
Coscoroba swan, (Coscoroba coscoroba) family with chicks, La Pampa, Argentina
Green-crowned brillian-t hummingbird (Heliodoxa jacula) visiting Torch ginger flower (Etlingera elatior), Poas Volcano National Park Costa Rica
Peruvian mantis (Oxyopsis peruviana) silhouette with sun behind, captive, occurs in South America
Western crowned pigeon, (Goura cristata), Aiduma Island, Triton Bay, near mainland New Guinea, Western Papua, Indonesian New Guinea
Karst islands in Misool archipelago, Raja Ampat, Western Papua, Indonesian New Guinea. December 2016
Wilsons bird-of-paradise (Cicinnurus respublica), Waigeo, Raja Ampat, Western Papua, Indonesian New Guinea
Scarlet Macaw (Ara macao) preening tail feather, Pantanal, Brazil
Alpine chamois (Rupicapra rupicapra rupicapra) in winter landscape with snow covered trees, Valsavarenche, Gran Paradiso National Park, Italy
Salar de Uyuni salt flat at sunset, Altiplano, Bolivia
Dingo (Canis lupus dingo) fighting on a beach. Fraser Island UNESCO World Heritage Site. Queensland, Australia, November
Hawks Head lookout over the Murchison River gorge, Kalbarri National Park, Western Australia, December 2015
Pika (Ochotona princeps) gathering plants to store for winter, Bridger National Forest, Wyoming, USA. August
Wood Duck (Aix sponsa). Male in breeding plumage flying at sunset. Acadia National Park, Maine, USA
Two small pearl-bordered fritillary butterflies (Boloria selene) resting on bluebell, Marsland mouth, North Devon, UK. May 2017
Kilchurn Castle, sunrise, early morning mist and light, a ruin on a rocky peninsula, the northeastern end of Loch Awe, in Argyll and Bute, Scotland, UK. March 2017
Treehopper (Alchisme grossa) Mindo, Pichincha, Ecuador
Brown Bear (Ursus arctos) adult portait, Finland, June
Broad-nosed weevil (Compsus sp. ) about to take off with wing elytra raised and wings extendedBroad-nosed weevil (Compsus sp.) about to take off with wing elytra raised and wings extended, Yasuni National Park, Orellana, Ecuador
Horned adder (Bitis caudalis) Swakopmund, Erongo, Namibia
Parsons Chameleon (Calumma parsonii) close up of eye and skin, Andasibe-Mantadia National Park, Moramanga, Madagascar
Clarion Burrowing Owl (Athene cunicularia rostrata), Clarion Island, Revillagigedo Archipelago Biosphere Reserve / Archipielago de Revillagigedo UNESCO Natural World Heritage Site (Socorro Islands)
Badger (Meles meles) adult in pine forest at sunset, Derbyshire, UK, June. Highly commended in the Habitat category of the British Wildlife Photography Awards (BWPA) Competition 2017
Female chalkhill blue butterfly (Lysandra coridon) with wings closed resting on Devils-bit scabious (Succisa pratensis), Hatch Hill, Somerset, UK. August 2016
Close up of pattern of the cerata of a nudibranch (Janolus savinkini). Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Tropical West Pacific Ocean
Orange-finned anemonefish (Amphiprion chrysopterus) guarding their recently spawned red eggs on a rock next to the host anemone. Rock Islands, Palau, Micronesia. Tropical west Pacific Ocean
Crocus flower (Crocus sp) stigma - close up
Orange cup coral (Tubastrea coccinea) Malpelo Island National Park, UNESCO World Heritage Site, Colombia, East Pacific Ocean
Juvenile Andean mouse opossum (Marmosops impavidus) eating a moth, El Oro, Ecuador
Southern stingray (Dasyatis americana) swimming over sand in shallow water, split level photo with blue sky and clouds. The Sandbar, Grand Cayman, Cayman Islands. British West Indies. Caribbean Sea
Mirrored portrait of a Weedy scorpionfish (Rhinopias frondosa). Bitung, North Sulawesi, Indonesia. Lembeh Strait, Molucca Sea. Digitally manipulated (the face is reflected down mid-line)