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Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, June
African lion (Panthera leo) lioness leaping over water, reflection in water, Okavango Delta, Botswana, July 2007
Red fox (Vulpes vulpes) two foxes at den in deep snow, Kamchatka, Far east Russia, November
Brown bear (Ursus arctos) catching salmon in river, Kamchatka, Far east Russia, August
RF - Polar Bears (Ursus maritimus) males fighting, Churchill, Canada, November
Camargue horse, two, one with mouth open. Bouches du Rhone, France May
Two wild Mustang stallions fighting in the White Mountain Herd Area, Wyoming, USA. August
Wild Mustang horse rearing, Pryor mountains, Montana, USA. June
Icelandic horse with mouth open, Iceland
Irish Gypsy cob (Equus caballus) grey stallion and piebald mare grazing on rough pastureland on Hackpen Hill with a Beech clump (Fagus sylvaticus) in the background, The Ridgeway
Grey Irish Gypsy cob stallion (Equus caballus) grazing rough pastureland on Hackpen Hill with Oilseed rape (Brassica napus) and other arable crops in the background, The Ridgeway
Bay Azteca stallion (Andalusian and Quarter Horse cross) running onto beach from waves, Summerland Beach, Ojai, California, USA
Chinese cobra (Naja atra) in threat stance, Shek Pik, southwestern coast of Lantau Island, Hong Kong, China
Indo-Pacific humpback dolphin ( Sousa chinensis) surfacing, Tai O, Lantau Island, Hong Kong, China
Atlantic salmon (Salmo salar) portrait, migrating to spawn in river, Gaspe Peninsula, Quebec, Canada, October
Mangrove hummingbird (Amazilia boucardi) female feeding chicks in nest, Pacific coast mangroves area, Costa Rica, Endangered species
Northern gannet (Morus bassanus) pair, courtship display, Machias Seal Island, Bay of Fundy, New Brunswick, Canada, May
Minke whale (Baelanoptera acutorostrata) breaching, Bay of Fundy, New Brunswick, Canada, July
Harbour porpoise, (Phocoena phocoena) porpoising, Bay of Fundy, New Brunswick, Canada, August
Yellow-spotted salamander eggs (Ambystoma maculatum) showing symbiosis with green algae, New Brunswick, Canada, May 2015
Brown Bear (Ursus arctos) female and cubs playing amongst the cotton grass, Finland, June
Black bears (Ursus americanus) standing on back legs, fighting, Minnesota, USA, June
Galapagos penguin (Spheniscus mendiculus) incubating egg, Puerto Pajas, Isabela Island, Galapagos
Barn swallow (Hirundo rustica) feeding young, Monmouthshire, Wales, UK, July
Swallow-tailed gull (Creagrus furcatus) calling, Genovesa Island, Galapagos
Wood pigeon (Columba palumbus) pair preening one another, The Netherlands
Three spot damselfish (Dascyllus trimaculatus) with Sea anemone home, Yap, Micronesia
Jaguar (Panthera onca) killing Spectacled caiman (Caiman crocodilus) in Piquiri River, Pantanal Mato Grosso, Brazil
Striped mackerel (Rastrelliger kanagurta) with mouths wide open as they swim through the water, filtering zoopankton with their gill rakers. Marsa Shouna, Port Ghalib, Marsa Alam, Egypt. Red Sea
Bluestreak cleaner wrasse (Labroides dimidiatus) sticking its head into the mouth of a Red Sea fusilier (Caesio lunaris) during a cleaning session. Small Crack, Sha ab Mahmud, Sinai, Egypt. Red Sea
Cheetah (Acinonyx jubatus) hunting Springbok (Antidorcas marsupialis) trying to trip up the prey, Kgalagadi, South Africa
Cheetah (Acinonyx jubatus) female and 8 / 9 week cubs, Masai Mara Game Reserve, KenyaCheetah (Acinonyx jubatus) female and 8/9 week cubs, Masai Mara Game Reserve, Kenya, September
Sawfly larvae (Nematus pavidus) defensive posture, Peatlands Park, County Armagh, Northern Ireland. September
Land crab (Geosesarma sp. ) which raids Pitcher plant (Nepenthes ampullaria) for preyLand crab (Geosesarma sp.) which raids Pitcher plant (Nepenthes ampullaria) for prey, Sarawak, Borneo
Peacock mantis (Pseudempusa pinnapavonis) in defensive posture; captive occurs in Burma and Thailand
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, Lake Victoria, Uganda
Calabar burrowing boa snake (Calabaria reinhardtii) in defensive ball, captive, occurs equatorial rain forest of West and central Africa. Head and tail are very similar
Trumpetfish (Aulostomus maculatus) using a Spanish Hogfish (Bodianus rufus) as a stalking horse. Spanish Hogfish is eating an arm of a starfish. Bonaire, Leeward Antilles, Caribbean
Pampas cat (Leopardus colocola) La Pampa Province, Argentina
Veined octopus (Amphioctopus marginatus) between clam shell halves. Ambon, Maluku Archipelago, Indonesia. Banda Sea, tropical west Pacific Ocean
Tawny nurse shark (Nebrius ferrugineus) group, individuals circling each other at dusk. Vavuu Atoll, Maldives. Indian Ocean
Northern goshawk (Accipiter gentilis) flying with squirrel prey, Finland. March
Blue tit (Cyanistes caeruleus) feeding young in the nestbox, Bavaria, Germany, May
Pheasant (Phasianus colchicus) male calling, East Frisian Islands, Wittbulten National Park. Wadden Sea UNESCO World Heritage Site, Germany, June
Lion (Panthera leo), females hunting buffalo, Masai-Mara Game Reserve, Kenya
Yellow-bellied toad (Bombina variegata) in defensive posture showing warning colours, on dried soil with mud cracks, Lower Saxony, Germany. August
Samoyed dog running in in snow, Ledyard, Connecticut, USA
Tiger (Panthera tigris), rolling on back and relaxing after feeding, Ranthambhore National Park, Rajasthan, India