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European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July
RF- Fluffy black kitten, age 10 weeks
Bottlenose dolphin (Tursiops truncatus) porpoising, Sado Estuary, Portugal. October
Frill-necked lizard (Chlamydosaurus kingii), with collar spread in threat display. Alice Springs, Northern territory, Australia
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, lake Victoria, Uganda
Brown capuchin monkey (Sapajus macrocephalus). displaying submissive behavior. Captive, Peru
Portrait of a meerkat (Suricata suricatta) or suricate, looking at the camera and standing to survey its territory on the edge of Makgadikgadi Pans National Park, Botswana, April
Armadillo girdled lizard (Cordylus cataphractus) coiled up in defensive posture in persons hand. Elands Bay, South Africa
Sable-and-white Border Collie puppy, age 8 weeks, in play-bow
Cavalier King Charles Spaniel x Poodle Cavapoo and red toy Poodle
Frill-neck Lizard (Chlamydosaurus kingii), displays on a termite mound. Northern Territory, Australia
Fosa / Fossa (Cryptoprocta ferox) male snarling, dry deciduous forest, Kirindy Forest, Western Madagascar, IUCN vulnerable species
Andalusian horsea, two stallions coming together in arena one dappled grey and one pure white. Spain
Iberian lynx (Lynx pardinus) walking on frosty grass, Parque Natural Sierra de Andujar, Andalucia, Spain. January
Rabbit (Oryctolagus cunniculus) juvenile grooming ears, Wiltshire, UK, May. Sequence 1 of 3
Yellowfin tuna (Thunnus albacares) hunting mackerel (Scomber colias) Tenerife, Canary Islands
Portrait of a wild adult female Bobcat (Lynx rufus) with Hispid cotton rat (Sigmodon hispidus) prey, Texas, USA. September
Portrait of a wild female Bobcat (Lynx rufus) kitten playing, Texas, USA. September
Tokay gecko (Gekko gecko) enacting a defensive display towards a perceived threat, Phuket Island, Thailand, March. Controlled conditions
Magnificent riflebird (Ptiloris magnificus alberti) male, on his display perch performing a display. Piccaninny Plains Sanctuary, Cape York Peninsula, Queensland, Australia
RF-Asian elephant (Elephas maximus) dust bathing. Jim Corbett National Park, Uttarakhand, India. (This image may be licensed either as rights managed or royalty free)
Yellow spotted monitor standing threat display {Varanus panoptes panoptes} Queensland, Australia
Lion (Panthera leo) males mock fighting / play fighting. Ngorongoro Crater, Ngorongoro Conservation Area (NCA), Tanzania
Southern right whale (Eubalaena australis) diving, with tail fluke splash. Monumento Natural Ballena Franca Austral, UNESCO World Heritage Site, Valdes Peninsula, Patagonia, Argentina
African lion (Panthera leo) and lioness after mating, Masai Mara National Reserve, Kenya
Lady Amhersts pheasant (Chrysolophus amherstiae) male displaying, Kanding, Sichuan, China. April
European rabbits (Oryctolagus cuniculus) fighting each other, Kiskunsag National Park, Hungary. June. Winner, 2019 Cadiz Photo Nature Competition
Polar bears (Ursus maritimus) fighting in water. Beaufort Sea, Kaktovik, Alaska, USA. Highly commended in the Polar Passion category of the Natures Best Photography Competition 2019
Bee-eater (Merops apiaster) male offering wasp as nuptial gift, Hungary. June
Timber rattlesnake (Crotalus horridus) females and newborn young at maternity site. Pennsylvania, USA. August
Fosa or Fossa (Cryptoprocta ferox)(sometimes erroneously fossa) male in dry deciduous forest. Kirindy, western Madagascar
Guanacos (Lama guanicoe), two males fighting, trying to over power opponent and bite each others testicles. Torres del Paine National Park, Patagonia, Chile. December
Bush cricket / Katydid (Tettigoniidae ), female leaf mimic ovipositing into branch. Manu Biosphere Reserve, Peru
RF- Wild male Jaguar (Panthera onca palustris) running through shallows of backwater of Cuiaba River in late afternoon sunlight. Northern Pantanal, Brazil
Giant River Otter (Pteronura brasiliensis) feeding on Striped Catfish or Cachara (Pseudoplatystoma fasciatum) amongst water hyacinth. Tributary off the Cuiaba River, Mato Grosso, Pantanal, Brazil
Net-casting / Ogre-faced Spider (Deinopis sp. ) with net. Active at night, Masoala National ParkNet-casting / Ogre-faced Spider (Deinopis sp.) with net. Active at night, Masoala National Park, Madagascar
Malayan colugo / Flying lemur (Cynocephalus variegatus) feeding on algae on tree bark. Danum Valley, Sabah, Borneo, Malaysia
North Pacific humpback whale (Megaptera novaeangliae kuzira) breaching at sunset, Kupreanof Island, Frederick Sound, Alaska, USA, July
Humpback whales (Megaptera novaeangliae) engaged in courtship, with the male hovering above the female. Vava'u, Tonga, Pacific Ocean
Twinspot snapper (Lutjanus bohar) shoal rushing up to spawn, releasing a cloud of milky white sperm and eggs in the water column. Caroline Islands, Palau, November
Ocean sunfish (Mola mola) and Pilot fish (Naucrates sp). Tenerife, Canary Islands
Bluestreak cleaner wrasse (Labroides dimidiatus) cleans among the sharp teeth of a predatory roving coral grouper (Plectropomus pessuliferus) beneath an overhang on a coral reef
Ruff (Philomachus pugnax) males displaying and fighting each other at a lek, Finland. May
Red fox (Vulpes vulpes) in mid air, snow diving / pouncing whilst hunting for rodents. Hayden Valley, Yellowstone National Park, USA. February
Little red flying fox (Pteropus scapulatus) female and suckling baby in flight, searching for place to roost. Nitmiluk National Park, Northern Territory, Australia
Cape buffalo (Syncerus caffer caffer) portrait, Red-billed oxpeckers (Buphagus erythrorhynchus) removing parasites. Nakuru, Kenya
Turtle dove (Streptopelia turtur) female feeding chicks on nest. Ranmore Common, near Dorking Surrey, England, UK. July
Fishing cat (Prionailurus viverrinus) walking with fish in mouth. Andhra Pradesh, India. Camera trap image. Photo© Anjani Kumar/Felis Images
