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Central American spider monkey (Ateles geoffroyi) juveniles hanging by tails and playing, Calakmul Biosphere Reserve, Yucatan Peninsula, Mexico, August
Sphinx hawk moth (Hemeroplanes triptolemus) caterpillar, snake mimic species, Amarakaeri Communal Reserve, Maijuna, Rio Napo, Peru
Harbour porpoises (Phocoena phocoena) - rare picture of small group Bay of Fundy, New Brunswick, Canada
Harbour porpoise (Phocoena phocoena) surfacing Bay of Fundy, New Brunswick, Canada
Peruvian Tarantula (Pamphobeteus sp. ) adult, walking over Humming Frog (ChiasmocleisPeruvian Tarantula (Pamphobeteus sp.) adult, walking over Humming Frog (Chiasmocleis royi) without preying on it. Los Amigos Biological Station, Madre de Dios, Amazonia, Peru
Red-footed booby (Sula sula), pair looking at one another on nest in tree. Egg is nest. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair perched in tree. Genovesa Island, Galapagos
Red-footed booby (Sula sula), pair nest building in tree. Darwin Bay, Genovesa Island, Galapagos
Nazca booby (Sula granti), pair billing. Genovesa Island, Galapagos. May 2015
Blue-footed booby (Sula nebouxii), six diving into sea. Espumilla Beach, Santiago Island, Galapagos
Jaguar (Panthera onca) female running along a sand spit chasing after a caiman. Northern Pantanal Cuiaba River, Mato Grosso, Brazil
Burrowing owl (Athene cunicularia) doing rain dance, spreading wings and dancing while it rained, Arizona, USA
Red-collared lorikeet (Trichoglossus rubritorquis) pair examining hole in Eucalyptus as a potential nesting hollow. Katherine, Northern Territory, Australia
Great bowerbird (Chlamydera nuchalis), male showing off treasures whilst female inspects bower. Lake Argyle, Kununurra, Western Australia
Kestrel (Falco tinnunculus) fighting Hungary, April
Common Snipe (Gallinago gallinago) autumn courtship display, Germany, August
Green iguana (Iguana iguana) rear view of basking in sun, Banco Chinchorro Biosphere Reserve, Caribbean region, Mexico, May
Thresher shark (Alopias pelagicus) swimming over seabed to be cleaned by cleaner wrasses, Cebu, Malaspascua, Philippines, September
Capercaillie (Tetrao urogallus) male displaying in rain, Kuusamo, Finland. March
Atlantic puffin (Fratercula arctica) portrait with beak open / calling. Shetland IslandsAtlantic puffin (Fratercula arctica) portrait with beak open/calling. Shetland Islands, Scotland, July
Pine grosbeak (Pinicola enucleator), male feeding juvenile, Finland, July
Black grouse (Lyrurus tetrix), male at lek, Finland, April
Eurasian hoopoe (Upupa epops) at nest in stone wall with hungry chicks gaping, Vendee, France, June
Commensal emperor shrimp (Periclimenes imperator) hitching a ride on a Nudibranch (Risbecia tryoni). Lembeh Strait, North Sulawesi, Indonesia
Grass snake(Natrix natrix) coiled round eggs, Alsace, France
Scarlet ibis (Eudocimus ruber), two birds perched on bushes, Coro, Venezuela. Small repro only
Young European eel (Anguilla anguilla) elvers, or glass eels, caught during their annual migration up rivers from the Bristol channel, swimming in a large holding tank at UK Glass Eels
Zebra (Equus quagga) leaping during stampede, Serengeti, Tanzania. Vignette added and right edge expanded
Nile crocodile (Crocodylus niloticus) Chobe National Park, Botswana, June
Giant panda (Ailuropoda melanoleuca) female, Huan Huan, holding baby age three months, Beauval Zoo, France, November 2017
Whale headed / Shoebill (Balaeniceps rex) vocalising. Swamps of Mabamba, Lake Victoria, Uganda
Norfolk Terrier dog, Otto, age 1 year
Hoffmanns Two-toed sloth (Choloepus hoffmanni) mother and baby, aged 2 months, in tree, Costa Rica. Rescued and released by Aviarios Sloth Sanctuary
Proboscis monkey (Nasalis larvatus) mother suckling infant, Sabah, Malaysia
Giant Manta (Manta birostris) with Remoras (Remora brachyptera) attached to both wings, offshore Santa Maria Island, Azores, Atlantic Ocean
African wild dogs (Lycaon pictus) play fighting, Mkuze, South Africa
Indonesian blue-tongued skink (Tiliqua gigas) with tongue extended, captive, from Papua New Guinea
Miniature American shepherd puppy in play-bow
Pilot whales (Globicephala macrorhynchus) just below surface, Tenerife, Canary Islands, Atlantic Ocean
Sperm whale (Physeter macrocephalus) resting just beneath surface, Faial Island, Azores, Portugal, Atlantic Ocean July
Short-finned pilot whale (Globicephala macrorhynchus) surfacing, South Tenerife, Canary Islands, Atlantic Ocean, July
Type 1 North-east Atlantic Killer whale / Orca (Orcinus orca) surfacing, Snaefellsnes Peninsua, western Iceland, January
Killer whale (Orcinus orca) diving and hunting for herring fish (Clupea harengus), Andenes, Andoya island, North Atlantic Ocean, Norway, April
Killer whale (Orcinus orca) tail slapping at sunset, transient race, Vancouver Island, British Columbia, Canada, July
Bowhead Whale (Balaena mysticetus) rubbing off flaking skin on the ocean bottom, east coast Baffin island, Qikiqtarjuaq, Nunavut, Canada, May
Dalmatian pelicans (Pelecanus crispus) underwater view of group feeding, Lake Kirkini, Greece. Highly commended in the GDT competition 2013
Jersey tiger moth (Euplagia quadripunctaria) large numbers resting on trunk of Sweetgum tree (Liquidambar orientalis) in the Petaloudes Valley, Rhodes island, Greece, August 2011
A Florida manatee (Trichechus manatus latirostrus) in upright posture with tail on river bed, outside Three Sisters Spring. Crystal River, Florida, USA