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Andean cock-of-the-rock (Rupicola peruvianus), two males perched in tree at lek in mid-altitude montane rainforest. Manu Biosphere Reserve, Amazonia, Peru
Parson's chameleon (Calumma parsonii), male hunting in forest canopy. Mitsinjo Forest, Andasibe-Mantadia National Park, eastern Madagascar
Bichon frise beside Colorado blue columbine (Aquilegia coerulea) flowers. Old Lyme, Connecticut, USA. May
Bichon frise standing on rock amongst Rhododendron flowers. Haddam, Connecticut, USA. May
RF - Andean cock-of-the-rock (Rupicola peruvianus), male perched in tree at a lek. Mid-altitude montane rainforest, Manu Biosphere Reserve, Amazonia, Peru
Blackbird (Turdus merula), male singing whilst perched in Ash (Fraxinus excelsior) tree, near Bath, England, UK. May
Free range Domestic pig (Sus scrofa domesticus) piglets play fighting, UK, August 2010
Broadclub cuttlefish hunting technique - mesmorises fish by pulsating colour along body
RF - Playful Silver tabby kitten rolling on back
White tailed sea eagle (Haliaeetus albicilla) perched on tree stump with fish. Hooded crow (Corvus cornix) flying in to attack in background. Danube Delta, Romania. May
Bee-eater (Merops apiaster) with Bumblebee prey, Sierra de Grazalema Natural Park, southern Spain, July
Whip scorpion (Heterophrynus elephas) hunting for food on a large tree root of the rainforest. Los Amigos Biological Station, Peru
Stag beetle (Lucanus cervus) males fighting on oak tree branch, Elbe, Germany, June
Black spider monkey (Ateles chamek) calling, captive, Peru
Red howler monkey (Alouatta seniculus) howling, captive, digitally modified
Leafcutter ant (Atta sp) carrying leaf along branch, Amazonian Rainforest, Yavari Valley, Loreto, Peru
Mozambique Spitting Cobra (Naja mossambica) ejecting venom Kruger, South Africa, Controlled conditions
Indo-Pacific humpback dolphin (Sousa chinensis) with fishing net damage around throat, Pearl River Estuary, Lantau Island, Hong Kong, China
RF - White lion (Panthera leo) male, portrait of head. Captive, Netherlands
RF - Meerkat (Suricata suricatta) resting on cool sand, Kgalagadi Transfrontier Park, South Africa
Straw-coloured fruit bat (Eidolon helvum), female flying carrying pup on front. Lamin, Gambia
Yacare caiman (Caiman yacare) with butterfly on snout, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Yacare caiman (Caiman yacare) on river bank, Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Jaguar (Panthera onca) male, hunting Capybara (Hydrochoerus hydrochaeris). The capybara while jumping away has cut the nose of the Jaguar with its toemail
Jaguar (Panthera onca) male, chasing a Caiman. Cuiaba River, Pantanal Matogrossense National Park, Pantanal, Brazil
Amur leopard snarling {Panthera pardus orientalis} captive
African lion (Panthera leo) growling at potential danger in the water (Panthera leo) Okavango Delta, Botswana
Lioness (Panthera leo) resting up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
Lion (Panthera leo) cub up a tree - only three populations of lions are known to do this habitually, Ishasha Sector, Queen Elizabeth NP, Uganda
African lion (Panthera leo) three subadults resting on the road, one snarling, Central Kalahari Game Reserve, Botswana
African lion (Panthera leo) young male stalking towards camera, Etosha National Park, Namibia October
Black Rhinoceros (Diceros bicornis) running. Etosha National Park, Namibia, January
RF- Jaguar head profle portrait (Panthera onca) head portrait captive. Pantanal, Brazil
Mountain gorillas (Gorilla gorilla beringei) grooming, members of the Munyaga group, Virunga National Park, North Kivu, Democratic Republic of Congo, Africa, Critically endangered
Mountain gorilla (Gorilla gorilla berengei) infant sitting on mothers head while she feeds on vegetation, Virunga Volcanoes National Park, Rwanda
Female Mountain gorilla (Gorilla beringei) carrying baby on her back, Volcanoes National Park, Rwanda, Africa
Wild male Snow Leopard (Panthera uncia) walking through barren terrain with snow covered patches, Altai mountains, Mongolia. February
Polar bear (Ursus maritimus) sow jumping while hunting for seals on sea ice, off the coast of Svalbard, Norway
Tabby cats, Picasso and Smudge, age 3 months, together. Smudge with his paws up on Picasso
Northern goshawk (Accipiter gentilis) perched on snag with grouse prey, Finland, March. March
Western capercaillie (Tetrao urogallus) Tver, Russia. May
Black Grouse (Tetrao tetrix) displaying at a lek in snow, Tver, Russia. April
Arctic fox (Vulpes lagopus) with Snow goose egg in mouth, mid moult from winter to summer fur, Wrangel Island, Far Eastern Russia, June
African lion (Panthera leo) lioness leaping over water, reflection in water, Okavango Delta, Botswana, July 2007
Red fox (Vulpes vulpes) two foxes at den in deep snow, Kamchatka, Far east Russia, November
Brown bear (Ursus arctos) catching salmon in river, Kamchatka, Far east Russia, August
RF - Polar Bears (Ursus maritimus) males fighting, Churchill, Canada, November
Camargue horse, two, one with mouth open. Bouches du Rhone, France May