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Two Arctic hares (Lepus arcticus) fighting, Northeast Greenland National Park, Greenland. Veolia Environnement Wildlife Photographer of the Year 2009: Mammal Behaviour category: Runner-up
Lioness (Panthera leo) carrying her cub, Masai-Mara Game Reserve, Kenya
Badger (Meles meles) adult, backlit at sunset, scratching its ears, Derbyshire, UK, May. Winner of the Animal Behaviour category of the British Wildlife Photography Awards (BWPA) Competition 2017
Atlantic Sailfish (Istiophorus albicans) attacking school of sardine (Sardinella aurita) bait ball. Isla Mujeres, Caribbean Sea, Mexico
Suffolk Punch heavy horse in field resting head anothers back, UK, September
Deepsea cirrate octopod {Stauroteuthis syrtensis} 830m, Gulf of Maine, Atlantic
Coastal brown bear (Ursus arctos) and cub, Lake Clarke National Park, Alaska, September
Two Giant pandas {Ailuropoda melanoleuca} climbing tree trunk in snow storm, Sichuan, China, captive
Polar bear cub 3 months (Ursus maritimus) playing in the front of the day den in march. Wapusk National Park. Churchill, Manitoba, Canada
Lion (Panthera leo) cubs playing, Masai-Mara Game Reserve, Kenya. Vulnerable species
Red Fox (Vulpes vulpes) walking along a fallen trunk in misty forest. Black Forest, Germany, November
Musk lorikeet (Glossopsitta concinna) feeding on flowers in a Eucalyptus tree, south-east Tasmania, Australia, February
Wren (Troglodytes troglodytes) singing, early morning in spring, Broxwater, Cornwall, UK. April
Herds of White-bearded wildebeest (Connochaetes taurinus albojubatus) crossing the Mara River. Northern Serengeti, Serengeti National Park, Tanzania
Goshawk (Accipiter gentilis) plucking a Willow Grouse (Lagopus lagopus) Liminka, Finland, March
Scottish wildcat (Felis silvestris grampia) captive, endemic to Scotland. September
RF - Australian Labradoodle, pouncing playfully
Brown bear (Ursus arctos) portrait in the Ozernaya River, Kuril Lake, South Kamchatka Sanctuary, Russia
Coconut / Veined octopus (Amphioctopus marginatus) hunts in the sand at night, while illuminated by a divers torch beam, Anilao, Batangas, Luzon, Philippines. Verde Island Passages, Pacific Ocean
Marine iguana (Amblyrhynchus cristatus) on shore, Punta Moreno, Isabela Island, Galapagos
Common dolphins (Delphinus delphis) surfacing, Fisheye lens. Pico, Azores, Portugal, June 2009
Hoopoe (Upupa epops) landing on branch, Hortobagy NP, Hungary
Hen harrier (Circus cyaneus) adult female landing on rock in moorland habitat, Glen Tanar Estate, Grampian, Scotland, UK, June
Red squirrel (Sciurus vulgaris) gnawing red deer antler for minerals, Cairngorms National Park, Highlands, Scotland, UK, June
King cobra (Ophiophagus hannah) in strike pose, Malaysia
Tree sparrow (Passer montanus) feeding a fledgling on a branch of Hawthorn (Crataegus monogyna) blossom, Perthshire, Scotland. June 2013
Kingfisher (Alcedo atthis) taking off from water with caught fish. Worcestershire, UK, March
Brandts cormorant (Phalacrocorax penicillatus) bursts through a school of Pacific chub mackerel (Scomber japonicus) while hunting beneath an oil rig
Common starling (Sturnus vulgaris) flocking prior to roosting, Scotland, January 2009
Boomslang (Dispholidus typus) Immature male hunting in thorn bush above deHoop Vlei, Western Cape, South Africa, December
Golden eagle (Aquila chrysaetus), Kuusamo, Finland, January
Australian sea lion (Neophoca cinerea) playful juvenile swimming upside-down, Carnac Island, Western Australia
Common loon (Gavia immer) nesting at edge of lake and calling, Acadia National Park, Maine, USA, August
Great crested grebe (Podiceps cristatus) pair performing courtship displaying at dawn, backlit and surrounded by mist, Cheshire, UK, April
Archerfish jets water at insect to dislodge it from branch {Toxotes chatareus} from SE Asia
Accumulation of land snails (Euryptyxis labiosa), escaping the desert heat by resting on a shrubs bark, Ayn Sahnawt, Sultanate of Oman, February
Shoebill stork (Balaeniceps rex) female feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, Lake Victoria, Uganda.. Sequence 2/13
Three banded armadillo {Tolypeutes tricinctus} in defensive ball. Captive, UK
European green crab (Carcinus maenas), an invasive species in North America, Kejimkujik Seaside National Park, Nova Scotia, Canada. July
RF- Fluffy black kitten, age 10 weeks
Bottlenose dolphin (Tursiops truncatus) porpoising, Sado Estuary, Portugal. October
Frill-necked lizard (Chlamydosaurus kingii), with collar spread in threat display. Alice Springs, Northern territory, Australia
Shoebill stork (Balaeniceps rex) feeding on a Spotted African lungfish (Protopterus dolloi) in the swamps of Mabamba, lake Victoria, Uganda
Brown capuchin monkey (Sapajus macrocephalus). displaying submissive behavior. Captive, Peru
Portrait of a meerkat (Suricata suricatta) or suricate, looking at the camera and standing to survey its territory on the edge of Makgadikgadi Pans National Park, Botswana, April
Armadillo girdled lizard (Cordylus cataphractus) coiled up in defensive posture in persons hand. Elands Bay, South Africa
Sable-and-white Border Collie puppy, age 8 weeks, in play-bow
Cavalier King Charles Spaniel x Poodle Cavapoo and red toy Poodle