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Great frigatebird (Fregata minor) in tree with pair of Nazca boobies (Sula granti) showing mating behaviour, Genovesa Island, Galapagos
Great spotted woodpecker (Dendrocopos major), male feeding juvenile in nest, Finland, June
Wolverine (Gulo gulo) walking in the snow. Wolverines paws are large, allowing them to walk easily through deep snow. Zapovednik, Kamchatka, Russia
Common eider duck (Somateria mollissima) female defending her nest from arctic fox (Vulpes lagopus). Longyearbyen, Spitsbergen. June
Australasian crested grebe pair (Podiceps cristatus australis) in courtship display. Ashburton Lakes, Canterbury, New Zealand. August
Great crested grebe (Podiceps cristatus) performing their weed dance during courtship. Den Oever, The Netherlands. July
Veined octopus (Amphioctopus marginatus) resting on top of the two halves of an old clam shell on the sandy seabed. These octopus regularly carry pieces of shell so they can use them as protective
Grayling (Thymallus) migrating to spawning in the Temnik River, Lake Baikal, Baikalsky Reserve, Siberia, Russia. May
Murmuration of Common starlings (Sturnus vulgaris) with Peregrine falcon (Falco peregrinus) hunting them, The Netherlands, November
Hummingbird Hawk-moth {Macroglossum stellatarum} adult in flight drinking nectar from lantana flower, Switzerland
Florida manatee (Trichechus manatus latirostris) swimming into a fresh water spring, Three Sisters Spring, Crystal River, Florida, USA
Mountain gorilla (Gorilla gorilla beringei) silverback Gihishamwotsi displaying, non group dominant, Sabyinyo Group, Volcanoes National Park, Virunga Mountains, Rwanda
Leopard (Panthera pardus) female fighting off male after he tries to mate with her. Londolozi Private Game Reserve, Sabi Sand Game Reserve, South Africa
Grey Andalusian stallion running on the beach at Ojai, California, USA
Grey whale (Eschrichtius robustus) at water surface with mouth open showing baleen plates, San Ignacio Lagoon, Baja California, Mexico
Great Bustard (Otis tarda) males displaying, Salisbury Plain, Wiltshire, UK, April
Patterns made in sand by Mint-sauce worms (Symsagittifera roscoffensis / Convoluta roscoffensis) sunbathing at low tide to allow the symbiotic algae to photosynthesise
Grizzly bear (Ursus arctos) catching a fish, Brooks Falls in Katmai National Park, Alaska, USA, July
Wood Duck (Aix sponsa), male flapping its wings, autumn colour reflected in water, Ohio, USA
Hen harrier (Circus cyaneus) female hovering over moorland, Glen Tanar Estate, Deeside, Scotland, UK, June
Cavalier King Charles Spaniels with tricolor and ruby colourations on beach, Texel, Netherlands
Jaguar (Panthera onca) male walking, Pantanal, Brazil
Arctic Fox (Alopex / Vulpes lagopus) standing on ridge, during moult from grey summer fur to winter white. Dovrefjell National Park, Norway, September
Lions (Panthera leo) - two brothers patrolling territorial boundary, affectionate behaviour, border of Serengeti / Ngorongoro Conservation Area (NCA) near Ndutu, Tanzania
Kingfisher (Alcedo atthis) perched with fish in beak. Worcestershire, UK, September
A male Leafy Seadragon (Phycodurus eques) carrying eggs. Wool Bay, Edithburgh, South Australia, Gulf of St Vincent, November
Polar bear (Ursus maritimus) juvenile shaking off snow after rolling around on the pack ice, arctic coast, 1002 area of the Arctic National Wildlife Refuge, Alaska, Beaufort Sea
Edwards Atlas Moth (Attacus edwardsii) in defensive posture, Bhutan, June
Ogre faced / Net-casting spider {Deinopis sp} with web held between legs that it will stretch over prey that walks below it. Masoala Peninsula National Park, north east Madagascar
Purebred grey Andalusian mare running in the snow, Longmont, Colorado, USA
Booted Racket-tail hummingbird (Ocreatus underwoodii) male flying, feeding from flower, Mindo, Ecuador, Andes, South America, January
Female Orang Utan (Pongo pygmaeus) [Sandy, born 29.04.82] sitting, holding two young. One [Samboja, born 09.06.05] is her own offspring, the other [Dayang
Sperm whale (Physeter macrocephalus) tail fluke above water as it dives below a rainbow, Dominica, Caribbean Sea, Atlantic Ocean, Vulnerable species. Digital composite
Common dolphin (Delphinus delphis) jumping, Pico, Azores, Portugal, June 2009
Rainbow lorikeets (Trichoglossus moluccanus) pair preening each other, perched in tree. Ulladulla, New South Wales, Australia
Silverback male Eastern lowland gorilla (Gorilla beringei graueri) intimidation display by chest beating, equatorial forest of Kahuzi Biega Park, Democratic Republic of Congo, Africa, March 2009
Bengal Tiger (Panthera tigris) six month old cub jumping on its mother, Bandhavgarh National Park, India
Andean cock-of-the-rock (Rupicola peruvianus), male displaying at lek in tree. Manu Biosphere Reserve, Amazonia, Peru
Mountain hares (Lepus timidus) boxing in snow, Scotland, UK, December
Spotted-tailed quoll (Dasyurus maculatus) scent marking in Monga National Park, New South Wales, Australia. Remote camera, triggered by movement
Brown hare, (Lepus europaeus), group of animals in field, Islay, Scotland, UK. March
Great Spotted Woodpecker (Dendrocopos major) adult male feeding young, Oisterwijk, The Netherlands, June
Piping Plover (Charadrius melodus) brooding three chicks with a fourth approaching, northern Massachusetts, USA.June. Endangered species
Great egret (Ardea alba) two fighting in flight, Lake Csaj, Kiskunsagi National Park, Pusztaszer, Hungary. January
Tabby kitten, 6 weeks, lying with head up and raised paw
Killer Whales (Orcinus orca) spy-hopping to observe Weddell Seal (Leptonychotes weddellii) in preparation to knock it from the ice by creating a wave. Marguerite Bay, Antarctic Peninsula, summer
Snow leopard mother (Uncia uncia) with cub, captive, occurs in mountains of central and southern Asia
Leaf cutter ants (Atta cephalotes) carrying sections of leaves, to be used for cultivating nutritious fungi, Santa Rita, Costa Rica
